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1.
Foam nests have evolved independently in several amphibian groups as an adaptive response to prevent predation and desiccation in dry environments. Nests are normally laid on ponds, or in underground galleries, humid forest leaf litter or terrestrial bromeliads. They are built when males or females beat a foam precursor associated with the egg masses extruded by the female. The spawning process requires the synchronic actions of the mating pair to obtain a hemispheric nest that protects the offspring. Herein, we describe the spawning behaviour of Engystomops pustulatus based on videos from 13 nesting couples from the lowlands of western Ecuador. Three variables were measured as indicators of male effort: duration of mixing events, duration of resting periods, and number of kicks per mixing event. We consider that not only male physical effort but also female behaviour influences nest structure. We suggest that nest building requires prolonged and intense physical activity by the male as well as the female’s steady position during spawning and female’s oviposition site selection. Nest building has two phases. In the first phase, the duration of resting periods, the duration of mixing events, and the number of kicks increase and are highly variable. During the second phase the three variables stabilise until the end. The volume of the nest increased mainly during the second phase. In four nesting events we observed kicking movements by the female. To our knowledge, this is the first time that female kicking has been observed in leptodactylid frogs. The function of this behaviour is unknown but our observations suggest that it may be triggered by insufficient male effort. Traditionally, female mate choice in Engystomops has been explained under models of indirect benefits exclusively. We argue that the prolonged male activity during nesting could influence female fitness directly. This will allow the operation of sexual selection via direct benefits. 相似文献
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Aloysius Domingo David Amar Karen Grütz Lillian V. Lee Raymond Rosales Norbert Brüggemann Roland Dominic Jamora Eva Cutiongco-dela Paz Arndt Rolfs Dirk Dressler Uwe Walter Dimitri Krainc Katja Lohmann Ron Shamir Christine Klein Ana Westenberger 《Cellular and molecular life sciences : CMLS》2016,73(16):3205-3215
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重组葡激酶(r-Sak)工程菌发酵工艺的研究 总被引:2,自引:0,他引:2
通过对影响重组葡激酶表达的几个主要因素的研究,建立了较为适宜的发酵条件:接种量4%;初始pH7.2;在A600nm=2.5时加20%蔗糖诱导,诱导后4~5h收获,发酵液上清r-Sak含量可达到250mg/L以上,酶活力可达到6500RU/ml以上。 相似文献
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R.M.罗斯 《国外科技新书评介》2007,(2):10-11
本书是编码理论教材,以具有重要理论和应用价值的纠错码为中心,特别着重于Reed—Solomon码及其派生码,给出这个理论的基本引论,在取材上兼顾了数学、计算机科学和电子工程等学科的不同需要,以初级的数学知识为起点,力求适应较广泛的读者群。 相似文献
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通过前臂试验对26种轻薄型羊毛机织精纺面料和7种其他纤维的轻薄型机织面料的刺痒感进行了比较研究,并用显微镜对部分轻薄型羊毛机织面料的表面纤维分布进行了研究。结果表明多数毛织物在温度为(24±1)℃、相对湿度为(65±5)%的条件下存在刺痒感,并且毛织物的刺痒感与其表面纤维的平面直径及直径大于26μm的表面纤维根数密切相关。 相似文献
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Swepson Pam Dick Bob Zuber-Skerrit Ortrun Passfield Ron Carroll Anne-Marie Wadsworth Yoland 《Systemic Practice and Action Research》2003,16(4):237-281
Action Learning, Action Research, and Process Management Association (ALARPM) is an organization of volunteers dedicated to the international expansion of action learning, action research, and process management, through world congresses. It has existed for over a dozen years now, despite significant stresses and strains, and has successfully conducted five world congresses with a sixth one in 2003. This history of ALARPM shows that a small group can set out to be international and inclusive from the beginning, so long as it also develops processes to sustain itself internally. 相似文献
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Arie Ron 《Cellular and molecular life sciences : CMLS》1974,30(9):1001-1002
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