Breeding ecology of White-faced Ibis ( Pleagadis chihi ) in the Upper Klamath Basin, California

The White-faced Ibis ( Plegadis chihi ) is a California state and U.S. Fish and Wildlife Service species of special management concern. We studied White-faced Ibis breeding ecology from May through July 1995 on Lower Klamath National Wildlife Refuge (NWR) in the upper Klamath Basin of California. A total of 2029 pairs nested in 3 colonies exclusively in early-successional hardstem bulrush ( Scirpus acutus ). Nest initiation dates ranged from 10 May to 12 June, and mean clutch size was 3.16. Reproductive success averaged 87% apparent nest success ( n = 115), 82% hatchability, 97% whole and partial brood survival, and 2.39 fledglings per successful nest. Mayfield estimates of nest survival were 79.1% during the laying and incubation period and 95% during the nestling period. Overall nest success as estimated by the Mayfield method was 75.4%. Our estimates of nest success are some of the highest reported anywhere in the literature for White-faced Ibis. Therefore, Lower Klamath NWR may maintain preferred White-faced Ibis breeding habitats in years of otherwise poor habitat conditions across the Intermountain West.     

Timing, distribution, and abundance of kokanees spawning in a Lake Tahoe tributary

We counted kokanee spawners and carcasses every 1-7 days from mid-September through mid-November in 1991 and 1992 in Taylor Creek, a tributary to Lake Tahoe, California-Nevada. Less than 1% of the spawning run entered Taylor Creek before flow from Fallen Leaf Lake was increased on 2 October 1991; in 1992 the peak occurred on 30 September or 1 October after flows increased on 29 September. In both years spawners concentrated in the middle three of five stream reaches below the impassable Fallen Leaf Lake dam. From tab-and-recovery experiments, the average longevity of male spawners in the stream was 3.5 days in 1991 and 2.8 days in 1992, whereas the average female longevity was 2.0 days in 1991 and 2.3 days in 1992. Observed carcasses accounted for less than 10% of spawners counted, suggesting removal by scavengers or high predation on prespawners. An estimated 1928 males and 1309 females spawned in 1991, and 8021 males and 8712 females spawned in 1992. Our estimate of 3237 spawners in 1991 compared favorably to our estimate of 3520 ± 1474 prespawners staging in Lake Tahoe in mid-September. An index of kokanee abundance in Lake Tahoe has historically been based on 1-day surveys every 1 November since 1960; however, estimated total spawner abundance was 19 times higher than the annual index of 158 spawners in 1991, 141 times higher than the index count of 100 spawners in 1992. The index count and mean fork lengths of spawners (278 ± 10 mm [2 SE] for males, and 248 ± 3 mm for females) in 1991 and 1992 were the lowest on record.     

Classification of the riparian vegetation along a 6-km reach of the Animas River, southwestern Colorado

Riparian ecosystems are important components of landscapes, particularly because of their role in biodiversity. A first step in using a ""coarse-filter"" approach to riparian biodiversity conservation is to determine the kinds of riparian ecosystems. These ecosystems vary substantially in plant species composition along a single river reach, as well as between rivers, and yet the river-reach scale has received little attention. We sampled the vascular plant composition of 67 contiguous patches of riparian vegetation along the reach of the Animas River, in southwestern Colorado's San Juan Mountains, that is relatively undisturbed by human land uses. Using cluster analysis and detrended correspondence analysis, we identified eight riparian community types along the reach. Using a new technique, we combined overstory size-class data and understory cover data to identify community types. The eight community types, which are in part the products of past floods, are spatially arranged along the reach in relation to variation in valley morphology, tributary location, and geomorphic landforms. These eight community types do not necessarily represent successional stages of a single potential vegetation type. This study at the river-reach scale suggests that sampling and analysis, as well as conservation, may need to be turned to the scale of patchiness produced by flood disturbances in the riverine landscape, since vegetation varies significantly at this scale.     

Sperm viability in the reproductive tract of females in a population of Sceloporus mucronatus exhibiting asynchronous reproduction

Asynchronous reproduction is a common phenomenon in high-elevation populations of lizards from Central México. Sperm storage in the reproductive tract of females is the mechanism for making oocyte fertilization possible. Our study addresses questions related to functional oviductal sperm storage of females mating on different dates throughout the reproductive season. A population of Sceloporus mucronatus with copulation in the summer and ovulation in the fall was chosen for this experiment. Eleven females that copulated in the field during June and 13 females that copulated in captivity during August were maintained in the laboratory until parturition. The number of pregnant females and the litter sizes produced in each experimental group were indicative of the viability of the stored sperm. Sperm stored in the reproductive tract of females were able to fertilize eggs after 4 months. No significant differences were found in the number of pregnant females between the 2 experimental groups nor in the litter sizes that they produced. We found that the amount of time sperm were held in the female reproductive tract (ca. 3 months) had no effect on the capacity of sperm to fertilize eggs. Histological examination of 8 oviducts collected before the mating season eliminated the possibility of sperm storage from one year to the next. In this system, sperm retention could have evolved as a response mechanism to deal with the asynchrony between sexes in the reproductive cycles. However, we cannot rule out alternative hypotheses.     

Distribution of the milliped Tylobolus utahensis Chamberlin, with remarks on T. fredericksoni (Causey) (Spirobolida: Spirobolidae)

Tylobolus utahensis Chamberlin, the only representative of the genus occurring in the southwestern deserts, ranges from central Inyo County, California, to the western periphery of Kane County, Utah. This distribution roughly corresponds to the northern limit of the Mojave Desert ecosystem and is also shown by the millipede Piedolus utus Chamberlin. (Spirobolida: Atopetholidae) and the centipede Theatops posticus (Say) (Scolopendromorpha: Cryptopidae). Tylobolus fredericksoni Causey, ostensibly from Douglas County, Kansas, is designated "" nomen dubium "" and disregarded pending collection of fresh material; Narceus gordanus (Chamberlin) is deleted from South Carolina and Tennessee. Tyobolus uncigerus (Wood) occurs north of the Columbia River in Klickitat County, Washington, and Hiltonius thebanus Chamberlin is referable to Onychelus Cook in the family Atopetholidae.     

Classification and ordination of the alpine plant communities of Railroad Ridge, White Cloud Peaks, Idaho

Alpine vegetation of Railroad Ridge in the White Cloud Peaks, Custer County, Idaho, was investigated to ascertain native communities and describe their distribution with respect to important environmental factors. Percent cover of all species and environmental information were recorded for 69 plots. Data were analyzed by TWINSPAN and CANOCO, which provided a classification and an ordination, respectively. Predominant communities are dominated by (1) Phlox pulvinata , (2) Carex elynoides , and (3) Ceum rossii , and other communities of lesser importance are dominated by (4) Artemisia campestris , (5) Eriogonum ovalifolium and Silene acaulis , and (6) Ivesia gordonii . Although the Artemisia campestris community occupies a relatively small portion of the study area, it is noteworthy because it is a community type not previously known for Idaho. The community co-dominated by Eriogonum ovalifolium and Silene acaulis is also unique and has not been documented in other Idaho alpine studies. Community distribution showed the strongest correlation with exposure, substrate type, and slope. In general, the study area supports fellfield communities that are representative of alpine areas in Idaho and some Great Basin mountain ranges, and dense turflike communities similar to ones found throughout western North America's temperate alpine regions.     

Winter macroinvertebrate communities in two montane Wyoming streams

Macroinvertebrate communities were examined on 4 winter dates over a 4-yr period in 2 high-altitude Rocky Mountain streams to document overwintering assemblages potentially experiencing spring acid pulses. Taxa richness values were comparable to other published lists for alpine and montane stream systems despite the fact that most literature reflected summer collections. Mean benthic density ranged from 1406 to 19,734 organisms/m 2 , and drift rates ranged from 0 to 1740 organisms/100 m 3 . Benthic collections showed higher taxa richness than drift collections while the Ephemeroptera and Plecoptera occurred in greater proportions in drift than in benthos. The Nemouridae (Plecoptera), Ephemerellidae and Heptageniidae (Ephemeroptera), Chironomidae (Diptera), and Hydracarina were the numerically dominant taxa in benthic collections. Grazer/scrapers and shredder/detritivores were always the numerically dominant functional feeding groups at all sites, composing 60-90% of the benthos. Predators, constituting approximately 15% of the community, occurred in the same relative proportion at all sites on all dates. Winter macroinvertebrate communities in these low-order, montane streams exhibit high taxonomic richness and benthic densities as great as lower-elevation mountain streams in the West.     

Habitat use and subspecific status of Merlins, Falco columbarius , wintering in central Utah

Most ornithological literature for Utah reports the status of the Merlin ( Falco columbarius ) as rare or scarce. Only the most recently published checklists (Behle and Perry 1975, Behle et al. 1985) record it as uncommon. Likewise, a 1998 printed checklist (Utah Ornithological Society 1998) list it as uncommon. Also misrepresented in the literature are the status and distribution of the 3 rather distinct subspecies, each of which occupies dissimilar habitats during the breeding season. We made random observations of Merlins in northern Utah beginning in the 1950s and then studied them continuously between 1992 and 1997 in Utah County, Utah. During the 1992-1997 period, we were able to clearly allocate 95 males and 76 females to subspecies, including within our count 58 falcons trapped (some fitted with radio telemetry). The winter habitat in which they occurred was categorized as either urban (mainly dense residential area) or rural (agricultural lands, dairy farms, or scattered homes). There was a statistically significant ( X 2 F. c. columbarius frequenting urban areas 68% of the time, and prairie parkland-breeding F. c. richardsonii frequenting rural habitats 78% of the time.     

Use of interstate highway overpasses and billboards for nesting by the common raven ( Corvus corax )

Common ravens are usually rather shy birds around their nests and avoid nesting too near human activity. We here report the use of overpasses along heavily traveled Interstate Highway 84 in Idaho and Utah for nest placement. Nests are within 6 m of passing vehicles.     

Vegetation characteristics of mountainous northeastern Nevada sagebrush community types

Vegetation characteristics of 15 sagebrush community types identified on the Humboldt National Forest, northeastern Nevada, are described. A total of 218 plant species were found over the 372 relatively undisturbed rangeland communities sampled. The dominant plant families encountered were the Asteraceae (45 taxa), Poaceae (32 taxa), Scrophulariaceae (14 taxa), and the Fabaceae (12 taxa). Average annual dry weight production of the community types ranged from about 400 kg/ha/yr on types with Artemisia nova as the dominant sagebrush species to 1,200 kg/ha/yr on some A. tridentata ssp. vaseyana community types. A general increase in species richness and vegetation plus litter ground cover was observed within community types as the dominant sagebrush species changed from A. nova to A. arbuscula to A. longiloba to A. tridentata spp. tridentata to A. tridentata ssp. vaseyana . Major differences in plant species production and constancy exist between the sagebrush community types studied.