Effects of browsing by native ungulates on the shrubs in big sagebrush communities in Yellowstone National Park

The effects of elk ( Cervus elaphus ), pronghorn ( Antilocapra americana ), and mule deer ( Odocoileus hemionus ) browsing on shrubs in big sagebrush ( Artemisia tridentata ) communities were monitored over a 31-year period in Yellowstone National Park. Ungulates were restricting Wyoming big sagebrush (spp. wyomingensis ) heights, size, and recruitment on the lower-elevation stratum only, while no such suppression was observed on the high-elevation stratum. Parallel increases in mountain big sagebrush (spp. vaseyana ) densities and cover occurred over the study period on both browsed and unbrowsed sites at the higher-elevation stratum, although big sagebrush, green rabbitbrush ( Chrysothamnus viscidiflorus ), and horsebrush ( Tetradymia canescens ) were slightly taller and crown sizes were slightly larger on unbrowsed than browsed sites. Wyoming big sagebrush utilization (percent leader use) was eight times higher (￣ x = 87 ± 7.2% by pronghorns, mule deer, and elk) on the low-elevation winter ranges stratum (the Boundary Line Area [BLA] portion of the winter range), while mostly mountain big sagebrush with leader use averaged only 11 ± 4.1% (nearly all by elk) on the high-elevation range stratum. In addition, annual aboveground biomass production of big sagebrush did not differ between browsed and unbrowsed study sites on the high-elevation stratum of the winter range. Population turnover was higher on browsed plots versus unbrowsed plots. No difference was observed in percent dieback of big sagebrush adult plants between browsed and unbrowsed plots at the higher stratum. Browsing did not influence the number of leaves or seedstalks per plant ( P > .05), but leaves averaged 45% longer and seedstalks 42% longer on browsed big sagebrush. Ungulate browsing, however, apparently suppressed production, germination, and survival of Wyoming big sagebrush on the low-elevation stratum. Numbers of Wyoming big sagebrush declined 43% and cover declined 29%, 1957-1990, on browsed sites on the BLA. Annual biomass production on browsed sites at the low-elevation stratum was only 6-35% that of unbrowsed sites, and big sagebrush recruitment was less on browsed sites. Percent leader use of big sagebrush did not differ between the period of ungulate reductions, 1962-1969, and the 1980s on the lower stratum (￣ x = 87% leader use), but utilization was less on higher portions of the winter range during the period of elk reductions (￣ x = 2%) than during the 1980s following cessation of elk controls (￣ x = 11%).     

Sprouting and seedling establishment in plains silver sagebrush ( Artemisia cana Pursh. ssp. cana )

The importance and nature of vegetative reproduction was compared with seedling establishment in plains silver sagebrush ( Artemisia cana Pursh. ssp. cana ). Sixty-three percent of plants excavated originated from rhizomes. Sites that experienced habitat disturbance did not have a significantly different number of plants originating from vegetative reproduction than did undisturbed sites. Parent rhizomes were significantly older than taproots, which were significantly older than aboveground stems. Rhizome systems were spread 3.3 times that of plant height. Seventy-nine percent of rhizomatous daughter plants were 100 cm or less from parent plants. Up to 52 sprouts were found on one rhizome. Seedling establishment was greatest during wet growing seasons, and vegetative reproduction was greatest during dry years.     

A sagebrush community type classification for mountainous northeastern Nevada rangelands

Vegetation characteristics of 15 sagebrush community types identified on the Humboldt National Forest, northeastern Nevada, are described. A total of 218 plant species were found over the 372 relatively undisturbed rangeland communities sampled. The dominant plant families encountered were the Asteraceae (45 taxa), Poaceae (32 taxa), Scrophulariaceae (14 taxa), and the Fabaceae (12 taxa). Average annual dry weight production of the community types ranged from about 400 kg/ha/yr on types with Artemisia nova as the dominant sagebrush species to 1,200 kg/ha/yr on some A. tridentata ssp. vaseyana community types. A general increase in species richness and vegetation plus litter ground cover was observed within community types as the dominant sagebrush species changed from A. nova to A. arbuscula to A. longiloba to A. tridentata spp. tridentata to A. tridentata ssp. vaseyana . Major differences in plant species production and constancy exist between the sagebrush community types studied.     

Vegetation characteristics of mountainous northeastern Nevada sagebrush community types

Vegetation characteristics of 15 sagebrush community types identified on the Humboldt National Forest, northeastern Nevada, are described. A total of 218 plant species were found over the 372 relatively undisturbed rangeland communities sampled. The dominant plant families encountered were the Asteraceae (45 taxa), Poaceae (32 taxa), Scrophulariaceae (14 taxa), and the Fabaceae (12 taxa). Average annual dry weight production of the community types ranged from about 400 kg/ha/yr on types with Artemisia nova as the dominant sagebrush species to 1,200 kg/ha/yr on some A. tridentata ssp. vaseyana community types. A general increase in species richness and vegetation plus litter ground cover was observed within community types as the dominant sagebrush species changed from A. nova to A. arbuscula to A. longiloba to A. tridentata spp. tridentata to A. tridentata ssp. vaseyana . Major differences in plant species production and constancy exist between the sagebrush community types studied.     

Suitability of shrub establishment on Wyoming mined lands reclaimed for wildlife habitat

Restoring coal mined land to pre-mining shrub cover, density, height, community composition, and diversity to renew wildlife habitat quality is a priority for reclamation specialists. Long-term shrub reestablishment success on reclaimed mined land in Wyoming and suitability of these lands for wildlife habitat are unknown. Fourteen reclaimed study sites, 10 yr old or older, were selected on 8 mines in Wyoming to evaluate shrub reestablishment and wildlife habitat value for antelope ( Antilocapra americana ) and sage grouse ( Centrocercus urophasianus ). Five sites were categorized as fourwing saltbush ( Atriplex canescens ) sites and 9 as fourwing saltbush/big sagebrush ( A. canescens/Artemisia tridentata spp. wyomingensis ) sites. Published data describing antelope and sage grouse-preferred habitat requirements in sagebrush-grassland steppe ecosystems were used to evaluate shrub community value of sampled sites for wildlife habitat. Mean shrub canopy cover, density, and height for fourwing saltbush sites were 5.8%, 0.23 m -2 , and 41.6 cm, respectively, compared to 5.6%, 0.61 m -2 , and 31.1 cm for fourwing saltbush/big sagebrush sites. Two fourwing saltbush and 4 fourwing saltbush/big sagebrush sites provided sufficient cover for antelope, while 2 fourwing saltbush and 4 fourwing saltbush/big sagebrush sites were adequate for sage grouse. Only 1 fourwing saltbush/big sagebrush site provided high enough shrub densities for sage grouse. One fourwing saltbush and 7 fourwing saltbush/big sagebrush sites provided ample shrub heights for antelope, while 1 fourwing saltbush and 8 fourwing saltbush/big sagebrush sites were sufficient for sage grouse. One fourwing saltbush and 1 fourwing saltbush/big sagebrush site provided enough grass, forb, and shrub composition for antelope, while no site in either reclamation type was satisfactory for sage grouse. Shrub diversity was 3 times higher for fourwing saltbush/big sagebrush sites (0.984) than for fourwing saltbush sites (0.328). Individually, sites seeded with multiple shrub species had higher canopy cover, density, and diversity compared with single-species shrub seedings. Achieving premining shrub cover, density, height, community composition, and diversity within existing bond-release time frames is unrealistic, considering that some native shrublands require 30-60 yr to reach maturity.     

Ant assemblages in intact big sagebrush and converted cheatgrass-dominated habitats in Tooele County, Utah

Biological invasions are one of the greatest threats to native species in natural ecological systems. One of the most successful invasive species is Bromus tectorum L. (cheatgrass), which is having marked impacts on native plant communities and ecosystem processes. However, we know little about the effects of this invasion on native animal species in the Intermountain West. Because ants have been used to detect ecological change associated with anthropogenic land use, they seem well suited for a preliminary evaluation of the consequences of cheatgrass-driven habitat conversion. In our study, we used pitfall traps to assess ant community assemblages in intact sagebrush and nearby cheatgrass-dominated vegetation. Ant abundance was about 10-fold greater in cheatgrass-dominated plots than in sagebrush plots. We also noted differences in diversity and evenness between habitat types at both the species and the functional-group levels of organization. At the species level, Shannon’s diversity index was greater in sagebrush plots than in cheatgrass-dominated plots. However, at the functional-group level, Simpson’s and Shannon’s diversity indices and the Brillouin evenness index were greater in cheatgrass-dominated plots than in sagebrush plots. Further, common species / functional groups tended to be more abundant while less common species / functional groups tended to be less abundant in cheatgrass-dominated plots compared to intact sagebrush plots. Patterns appear to be at least partially related to resource availabilities. This initial survey of ant communities from intact-native and altered vegetation types may be indicative of similar trends of biodiversity shifts throughout the Intermountain West where cheatgrass has successfully replaced native species. We also discuss the implications of ant communities on land management activities, specifically in the context of aridland restoration.     

Invasion of big sagebrush ( Artemisia tridentata ) by white fir ( Abies concolor ) on the southeastern slopes of the Warner Mountains, California

White fir ( Abies concolor ) appears to be invading vegetation dominated by big sagebrush ( Artemisia tridentata ) on the southeastern slopes of the Warner Mountains of extreme northeastern California. The time of initial tree establishment within the shrubs was determined by increment borings. Possible causes of the invasion involving specific climatic conditions, fire history, and grazing use of the area during years of the establishment were explored. Heavy grazing by domestic livestock, particularly sheep, appears responsible for altering the sagebrush - grass vegetation and allowing tree invasion.     

Short-term effects of a prescribed burn on songbirds and vegetation in mountain big sagebrush

I measured songbird abundance and vegetation cover in and around a 420-ha prescribed burn in a mountain big sagebrush community located at 2133 m elevation. Data were collected during the 3rd growing season after the fire. Brewer's Sparrow and Sage Thrasher occurred in lower abundance on sites that were largely or completely burned relative to sites that were outside the fire perimeter or within unburned islands of shrubs. The number of Brewer's Sparrow detections was linearly related to remaining sagebrush cover. In contrast, Horned Lark occurred at higher abundances on sites where shrub cover had been removed in the prescribed burn. Cover of perennial grasses and cover of 4 of the 5 most common forbs was greater on burned sites than on unburned sites.     

Vegetation response to prescribed fire in Dinosaur National Monument

Much of western North America is dominated by dense, monotypic, late seral stands of big sagebrush ( Artemisia tridentata Nutt.). These stands often have depauperate understories with limited species richness, diversity, and herbaceous cover. The National Park Service at Dinosaur National Monument, Colorado, is using both strategic and natural prescribed fire in Wyoming big sagebrush ( Artemisia tridentata ssp. wyomingensis Beetle and Young) communities to foster intra-community (α -scale) and landscape diversity. This study analyzed an accumulated foliar cover data set between paired burn and control areas on 6 different sites during the last 20 years. Across the monitoring period, mean total vegetation cover of all combined sites was 44% control and 42% burn. Total vegetation cover in burn areas was higher than or equal to paired control areas within 2-3 years post-burn. Shrubs were essentially eliminated in burn areas, but perennial grass cover was 10-35% higher. Mean number of species on all sites and years combined was 17 control and 18 burn. Species richness was different on only 1 site-year, Dry Woman 1995 ( P = 0.001, 15 control, 9 burn). Species similarity by site and between treatments ranged from 44% to 75%. Differences in Shannon-Weiner diversity index values between paired sites occurred in 6 of 20 years ( P < 0.05). Index value differences on these 6 sites were due to a large annual grass component in burn areas. Prescribed burning successfully shifted late successional sagebrushdominated communities to earlier herbaceous-dominated successional stages without lowering total vegetation cover, while maintaining -scale diversity and species richness.     

Summer habitat use by Columbian Sharp-tailed Grouse in western Idaho

We studied summer habitat use by Columbian Sharp-tailed Grouse ( Tympanuchus phasianellus columbianus ) in western Idaho during 1983-85. Vegetative and topographic measurements were recorded at 716 locations of 15 radio-tagged grouse and at 180 random sites within the major vegetation/cover types in the study area. The mean size of summer home ranges was 1.87 ± 1.14 km 2 . Of eight cover types identified in the study area, individual grouse used the big sagebrush Artemisia tridentata ) cover type more than or in proportion to availability, the low sagebrush ( A. arbuscula ) in proportion to availability, and avoided the shrubby eriogonum ( Eriogonum spp.) type. Characteristics of the big sagebrush cover type that Sharp-tailed Grouse preferred include moderate vegetative cover, high plant species diversity, and high structural diversity. Grouse used areas of dense cover (i.e., mountain shrub and riparian cover types) primarily for escape cover. Compared with random sites, grouse selected areas with (1) greater horizontal and vertical cover, (2) greater canopy coverage of forbs typically decreased by livestock grazing, (3) greater density and canopy coverage of arrowleaf balsamroot ( Balsamorhiza sagittata ), and (4) greater canopy coverage of bluebunch wheatgrass ( Agropyron spicatum ) in the big sagebrush cover type in 1984 and the low sagebrush cover type in 1985. The importance of the native perennials arrowleaf balsamroot and bluebunch wheatgrass became apparent during a drought year when many exotic annuals dried up and provided no cover. Overall, grouse selected vegetative communities that were least modified by livestock grazing.     

Use and selection of brood-rearing habitat by Sage Grouse in south central Washington

Sage Grouse ( Centrocercus urophasianus ) brood-habitat use was examined during 1992 and 1993 at the Yakima Training Center in Yakima and Kittitas counties, Washington. During the 2 yr we followed 30 broods, of which 12 persisted to 1 August (￣ x = approximately 1.5 chicks/brood). Food forb cover was greater at all brood locations than at random locations. Hens with broods in big sagebrush/bunchgrass habitat ( Artemisia tridentata/Agropyron spicatum ) selected for greater food forb cover, total forb cover, and lower shrub heights; broods in altered big sagebrush/bunchgrass habitats selected greater tall grass cover and vertical cover height; broods in grassland showed no preference for any measured vegetation characteristics. During the early rearing period (post-hatching-6 wk) each year, broods selected sagebrush/bunchgrass. Broods in 1993 made greater use of grasslands than in 1992 and selected grassland during the late brood-rearing period (7-12 wk). Broods selected for sagebrush/bunchgrass during the midday, but 52% of brood locations in the afternoon were in grassland. Tall grass cover was greater at morning (0500-1000 h) and afternoon (1501-2000 h) brood locations than at midday (1001-1500 h) and random locations. Midday brood locations had greater shrub cover and height than morning and afternoon locations. Selection of habitat components was similar to the results of other studies, but habitat conditions coupled with a possible lack of alternate brood-rearing cover types resulted in low survival of chicks.     

Reptile population changes with manipulation of Sonoran Desert shrub

The diversity and abundance of reptiles were studied in three vegetation types on the Santa Rita Experimental Range, Arizona. Total reptile sightings were greatest in undisturbed mesquite and mesquite with irregularly shaped clearings. No zebra-tailed lizards ( Callisaurus draconoides ) or desert spiny lizards ( Sceloporus magister ) were seen, and significantly fewer western whiptails ( Cnemidophorus tigris ) were in the mesquite-free area. Only the Sonora spotted whiptail ( Cnemidophorus sonorae ) was significantly more abundant in the mesquite-free area than in the undisturbed mesquite. In an effort to increase grass production for cattle in mesquite grasslands, it is preferable to clear irregularly shaped areas rather than to attempt total mesquite removal, if reptiles are to be considered.     

Winter habitat-use pattern of elk, mule deer, and mosse in southwestern Wyoming

Habitat-use patterns of mule deer, elk, and moose were determined on two winter range near Kemmerer, Wyoming. Mule deer used areas with the least snow depth and dominated by sagebrush. Elk were located more often than expected on wind-swept hills but used sagebrush communities more frequently as snow depths increased. Moose were generally found associated with broad, riparian zones. All three species occasionally used the same area but differed in their use of specific vegetation types and topography.     

Competition Between Adult and Seedling Shrubs of Ambrosia dumosa in the Mojave Desert, Nevada

Seeds of the perennial shrub Ambrosia dumosa germinated in abundance following 11 days of rain during August 1983 at a study site in the northern Mojave Desert. Seedling establishment, growth, and reproduction were observed in natural vegetation and in an area that had been previously cleared of vegetation. For 5,527 A. dumosa seedlings, percent survival in April 1986 averaged 3% in the undisturbed vegetation and 58% in the denuded area. Seedlings occupying the cleared area had grown to sizes up to 0. 1 m 3 m by October 1984; some produced flowers and fruit in the spring of 1985. Surviving seedlings in the undisturbed vegetation were all smaller than 0.001 m 3 and did not reproduce. These pronounced differences in growth, survival, and reproduction associated with the presence or absence of adult shrubs demonstrated an intense competition that is incompatible with indications of mild competition from nearest-neighbor analyses. I therefore hypothesize that competition for water occurred, not by competition for water in two dimensions but by rapid use of a common resource, as if several people were drinking with straws from a common cup. This temporal mechanism would strongly favor adults over seedlings.     

Seasonal microhabitat relationships of blue grouse in southeastern Idaho

Microhabitat characteristics of blue grouse ( Dendragapus obscurus ) were analyzed in breeding and wintering habitats in southeastern Idaho. Breeding habitats typically were open sagebrush ( Artemisia spp.), mixed shrub, mountain mahogany ( Cercocarpus ledifolius ) , and maple ( Acer grandidentatum ) stands on east to south facing aspects of slopes below 2100 m elevation. Breeding blue grouse selected areas with approximately a 50:50 or greater open to cover ratio. Blue grouse selected areas with higher tree coverage than available on average within the mixed shrub vegetation type. Hens with broods preferred sites with relatively tall (>50 cm) herbaceous vegetation. During autumn and winter, blue grouse preferred high elevation (>2285 m) stands of open (50% tree cover) conifer. Douglas-fir ( Pseudotsuga menziesii ) were preferred as winter roost trees. Sites selected in winter had significantly more Douglas-fir than those selected in autumn.&nbsp;     

Response of sagebrush steppe species to elevated CO 2 and soil temperature

Elevated atmospheric CO 2 may cause long-term changes in the productivity and species composition of the sagebrush steppe. Few studies, however, have evaluated the effects of increased CO 2 on growth and physiology of species important to this ecosystem. Since the response of plants to elevated CO 2 may be limited by environmental factors, soil temperature was also examined to determine if low soil temperatures limit CO 2 response. To determine how CO 2 and soil temperature affect the growth of species native to the sagebrush steppe, bottlebrush squirreltail [ Elymus elymoides (Raf.) Swezey], Thurber needlegrass ( Stipa thurberiana Piper), and Wyoming big sagebrush ( Artemisia tridentata ssp. wyomingensis Beetle) were grown in ambient (374 mL L -1 ) or high (567 mL L -1 ) CO 2 and low (13° C) or high (18° C) soil temperature for approximately 4 months. Although soil temperature affected the growth of squirreltail and needlegrass, temperature did not modify their response to elevated CO 2 . Total biomass of sagebrush was consistent across soil temperature and CO 2 treatments, reflecting its slow-growing strategy. All 3 species had higher leaf water-use efficiency at elevated CO 2 due to higher net photosynthesis and lower transpiration rates. We conclude that elevated CO 2 and soil warming may increase the growth of grasses more than shrubs. Field studies in the sagebrush steppe are necessary to determine if differences in biomass, resulting from changes in CO 2 and soil temperature, are exhibited in the field.     

Selection of big sagebrush by sage grouse

Feeding sites of wintering sage grouse ( Centrocercus urophasianus ) were located, one each in stands of three subspecies of big sagebrush ( Artemisia tridentata : ssp. tridentata , basin; ssp. vaseyana , mountain; and ssp. Wyomingensis , Wyoming). Evidences of differential use of plants within subspecies were observed. Whole leaves from fed-on and nonfed-on big sagebrush plants were examined for intrasubspecies chemical comparisons of crude protein, phosphorus, in vitro digestibility, and monoterpenoids. No significant differences were detected except for in vitro digestibility of Wyoming fed-on and nonfed-on big sagebrush and monoterpenoid content of basin big sagebrush. Nutritive content of all three subspecies was high, which may in part help to explain wintering sage grouse weight gains.     

Long-term effects of tebuthiuron on Bromus tectorum

Use of herbicides to thin dense stands of Artemisia spp. (sagebrush) can free up resources for herbaceous plants and increase forage production, but may also facilitate weed invasion. We revisited a sagebrush thinning experiment in a north central Wyoming big sagebrush–grassland 11 years after application of tebuthiuron (N-[5-(1,1- dimethylethyl)-1,3,4-thiadiazol-2-yl]-N-N′-dimethylurea) to determine the long-term responses of shrubs, available soil resources, perennial grasses, and Bromus tectorum L. (downy brome). Tebuthiuron reduced shrub cover by more than half, from 31% in untreated plots to 15% in treated plots ( P = 0.002), and increased downy brome cover approximately 4-fold, from 0.9% in untreated plots to 3.5% in treated plots ( P = 0.02). Treatment with tebuthiuron also resulted in marginally significant increases in cover of perennial grasses (from 9% to 12.3%; P = 0.07) and bare ground (from 39.1% to 43.9%; P = 0.08). In comparisons of resource availability among microsites, available NO 3 was higher under dead sagebrush than under live sagebrush ( P = 0.03). No significant differences in soil water content were detected. The relatively recent expansion of downy brome populations at this site and the high NO 3 –N levels observed under dead sagebrush suggest that conditions facilitating downy brome invasion may persist for many years following sagebrush thinning. We demonstrate that sagebrush thinning can cause increases in downy brome populations years after initial treatment and suggest that managers should use caution when considering thinning sagebrush if downy brome is present, even if initial populations are small.     

Ecological classification and monitoring model for the Wyoming big sagebrush shrubsteppe habitat type of northeastern Wyoming

The purposes of this study were to develop a multivariate statistical model related to plant succession, to classify by seral stage, and to monitor succession in Wyoming big sagebrush shrubsteppe habitat ( Artemisia tridentata ssp. wyomingensis / Agropyron smithii–Bouteloua gracilis ) in Thunder Basin National Grassland, Wyoming. This model can be used by range and wildlife managers to evaluate management alternatives by assessing changes in plant species cover and composition within and between seral stages. Four ecological seral stages that represent early to late succession were quantitatively identified with an estimated 92% accuracy. Three key plant species provided the necessary information to define seral stages and monitor trends. Percent canopy cover and percent frequency (used to calculate index values: % canopy cover × % frequency of occurrence) of Wyoming big sagebrush ( Artemisia tridentata ssp. wyomingensis ), western wheatgrass ( Agropyron smithii ), and blue grama ( Bouteloua gracilis ) are the only field measurements required for this model.     

Effects of 2,4-D on a Populus tremuloides community in the western United States-22 years after treatment

Normal 0 false false false EN-US X-NONE X-NONE MicrosoftInternetExplorer4 st1\:*{behavior:url(#ieooui) } /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Table Normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-priority:99; mso-style-qformat:yes; mso-style-parent:""; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:11.0pt; font-family:"Calibri","sans-serif"; mso-ascii-font-family:Calibri; mso-ascii-theme-font:minor-latin; mso-fareast-font-family:"Times New Roman"; mso-fareast-theme-font:minor-fareast; mso-hansi-font-family:Calibri; mso-hansi-theme-font:minor-latin; mso-bidi-font-family:"Times New Roman"; mso-bidi-theme-font:minor-bidi;} Quaking aspen ( Populus tremuloides Michx.) stands were accidentally sprayed with 2,4-D in a sagebrush control program in western Wyoming in 1958. We visited the site during the summer of 1981 to evaluate the long-term effect on the aspen trees and the associated vegetation. Initially, some observers believed that the aspen had been &ldquo;destroyed.&rdquo; Subsequent data indicate just the opposite&mdash;aspen stocking appeared to have been improved by the treatment. On two of the sampled clones, 22 years after spraying, there were approximately 17,000 more suckers/ha on the sprayed than on the unsprayed plots. These are adequate numbers to restock the site to pretreatment densities. Although undergrowth vegetation appeared to be changed as a result of the treatment, this cannot be attributed solely to the herbicide because heavy grazing, mostly by domestic livestock, has occurred on the grazing allotment. A similarity index was calculated between sprayed and unsprayed portions of the same aspen clones. Forbs were still less on the sprayed areas, whereas grasses were similar on the sprayed and unsprayed areas. Spraying apparently does not have as adverse an effect on aspen communities as some conservationists originally thought. &nbsp;&nbsp;