Response of winterfat ( Ceratoides lanata ) communities to release from grazing pressure

Sixteen study sites were established in grazed and ungrazed stands of winterfat in Kane County, Utah. The area is located within the winter range of cattle and along U.S. Highway 89 between Kanab, Utah, and Page, Arizona. Road construction in 1957 dissected several winterfat communities, and following fencing part of the communities were released from grazing. Differences in species composition, vegetation, and soil characteristics between grazed and ungrazed sites were assessed. Major differences in site characteristics appeared due to the influence of winter grazing by cattle. Winterfat and Indian ricegrass showed increased cover on the nongrazed sites following release from grazing pressure. Winterfat also showed significant negative interspecific association patterns with all major species.     

Grazing impacts on sagebrush communities of central Utah

Twenty-three study sites were established in the sagebrush communities bordering Utah Lake. Relic (ungrazed) and grazed stands were represented in the sample. Differences in species composition, vegetation, and soil characteristics were assessed. Major differences in species composition and vegetative characteristics were due to the influence of grazing. Major changes were loss of native perennial grass cover, and increases in cover from introduced annual species. Differences in the soil characteristics were due to habitat rather than grazing influences.     

Vegetation changes after livestock grazing exclusion and shrub control in the southern Chihuahuan Desert

Vegetation cover and production were evaluated after nearly 7 years of livestock grazing exclusion and shrub control in an area with a long history of heavy livestock grazing in the southern Chihuahuan Desert, Mexico. An exclosure was established to prevent livestock grazing. In half of the excluded area, the main shrub, Larrea tridentata , was mechanically controlled. Outside the exclosure, heavy livestock grazing occurred as customary and shrubs were not controlled. Absence of grazing resulted in 50% higher grass cover and 35% higher total biomass. Larrea tridentata cover was twice as high on the grazed area as on the ungrazed area. Vegetation cover was dominated by grasses (42%) in the ungrazed area, whereas in the grazed area, cover was equally divided between grasses (28%) and shrubs (27%). Shrub control did not affect vegetation cover or herbage production. Multivariate analysis confirmed that inside the excluded area, shrub control had little impact on the plant community. The effect of grazing, however, clearly distinguished the community outside the exclosure from that inside the exclosure.     

Grasshopper densities on grazed and ungrazed rangeland under drought conditions in southern Idaho

Low-density grasshopper populations were sampled at 15 pairs of rangeland sites in south central Idaho. One site of each pair had not been grazed by livestock for at least 10 years. Grazed sites were managed under normal grazing regimes established by the Bureau of Land Management. Mean grasshopper density was higher on ungrazed sites than on grazed sites. Proportions of Melanoplus sanguinipes were higher on ungrazed sites than on grazed sites and were higher on annual grasslands than on other vegetation types. Effects of grazing appeared to be independent of vegetation type. Proportions of Comhpocerinae, a subfamily of grasshoppers that feeds almost exclusively on grasses, were affected by vegetation type, but not grazing. Crested wheatgrass seedings supported the highest proportions of Gomphocerinae. Proportions of Oedipodinae were affected by grazing and vegetation type. Higher proportions of Oedipodinae were found on grazed sites than on ungrazed sites, and on sagebrush/grass sites than on annual grasslands. Results indicate that livestock grazing during drought conditions tends to reduce grasshopper populations in southern Idaho rangeland.     

Effects of grazing exclusions on rangeland vegetation and soils, east central Idaho

Nineteen exclosures on sagebrush steppe and shadscale rangelands, varying in age from 18 to 38 years, were sampled for plant species richness, plant composition, indicators of soil erosion, ground cover, vegetative cover, and herb-low shrub layer screening cover. Features within the exclosures were compared with adjacent sites of the same size that were open to grazing by livestock and wildlife. Species richness typically was slightly greater inside exclosures than in adjacent grazed sites (median = 2 more species inside exclosures), but the difference was not significant ( P = 0.16). Similarity of plant community composition between exclosures and adjacent grazed sites ranged from 45% to 82%. Evidences of soil movement, soil pedestals, and soil flow patterns were all more pronounced outside exclosures than inside ( P ≤ 0.02), even though many sites were on flat to mild slopes (median slope = 12%). Meta-analysis of the 19 exclosure sites indicated that grazing exclusion resulted in less bare ground cover compared with adjacent grazed sites ( P ≤ 0.05). The effect of grazing exclusion on visible soil surface cryptogams was significant ( P ≤ 0.05), with generally greater cover inside exclosures. Cryptogam cover differences between grazed sites and exclosures tended to increase with the number of years of grazing exclusion ( r = 0.64, P = 0.046). Pseudoroegneria spicata , a principal livestock forage, averaged greater basal cover inside exclosures than outside on 4 of 10 sites where it occurred, although no exclosure sites had greater P. spicata cover on grazed sites. Meta-analysis of the 10 sites indicated that grazing exclusion resulted in greater P. spicata cover compared with adjacent grazed areas ( P ≤ 0.05). Poa secunda , a short-growing grass that initiates growth early in the spring and is not important livestock forage, averaged greater basal cover outside exclosures on 5 of 15 sites where it occurred. Meta-analysis of the 15 sites indicated a significant treatment effect ( P ≤ 0.05), with greater Poa secunda basal cover outside exclosures. Grazing exclusion resulted in greater screening cover in the herb-low shrub layer (0-0.5 m height; P ≤ 0.05). These results indicate that despite improved livestock grazing management over the past half century, livestock grazing still can limit the potential of native plant communities in sagebrush steppe ecosystems, and that the health of semiarid ecosystems can improve with livestock exclusion in the absence of other disturbances. A few exclosure sites were similar for the measured parameters, suggesting that these sites were ecologically stable and that exclusion of livestock grazing was not sufficient to move succession toward more pristine conditions, at least within the time periods studied. Managed disturbance such as fire or mechanical brush treatments may be necessary to restore herb productivity on these ecologically stable sites.     

The effect of livestock grazing on the rainbow grasshopper: population differences and ecological correlates

This study examined the effect of livestock grazing on a common herbivore in semiarid grasslands of the Southwest: the rainbow grasshopper, Dactylotum variegatum . Population attributes and key environmental variables were compared between sites on active cattle ranches and sites on a 3160-ha ungrazed sanctuary. Although density of D. variegatum nymphs did not differ significantly between grazed and ungrazed sites, adult density was significantly lower on grazed sites, and sex ratios differed between grazed and ungrazed sites over time. Grazed sites had higher percentages of bare ground and fewer Baccharis pteronioides , a common shrub. However, only bare ground was highly correlated with adult D. variegatum density; lower adult densities were associated with more bare ground. At a smaller scale, D. variegatum tended to be more abundant in areas with B. pteronioides on grazed sites and less abundant in areas on ungrazed sites. This study suggests that cattle grazing influences ecological variables important in determining density, sex ratio, and spatial distribution of D. variegatum at both large and small spatial scales.     

C 3 and C 4 Species Changes Identified by δ 13 C values of soil organic matter in a Colorado prairie

We measured carbon isotope signatures (δ 13 C) from 0-10 cm and 10-20 cm soil depth intervals for grassland soils near Boulder, Colorado. These grasslands included tall-, short-, and mixed-grass prairies that were grazed, ungrazed, or hayed. Soils exhibited δ 13 C signatures consistent with observations that current sites are a mix of C 3 and C 4 species, with C 3 plants more abundant in mixed-grass than in native tall- or shortgrass prairies. The δ 13 C signatures were not significantly different for grassland types; however, management treatments (grazing, no grazing, haying) significantly influenced changes in soil δ 13 C signatures from the 0-10 cm to 10-20 cm soil depth intervals. We observed a correlation ( r = 0.63) between isotopic values of surface soils and percent native species in total vegetation cover. Overall, the community type with the lowest percentage of nonindigenous species cover had the most enriched δ 13 C signature. Sites currently grazed by prairie dogs, cattle, or both herbivores had stronger C 3 signatures, indicating that grazing may have increased C 3 plant productivity in these communities at the expense of C 4 grasses. This finding differs from studies of native shortgrass steppe where grazing has the opposite effect on the relative abundance of these 2 functional groups of plants. This result, along with the correlation between C 3 isotopic values and nonnative vegetation abundance, provides evidence that management practices that maintain dominance of C 4 grasses should be encouraged.     

Effects of long-term livestock grazing and habitat on understory vegetation

The herbaceous understory stratum contains most of the plant diversity in ponderosa pine ( Pinus ponderosa P. & C. Lawson var. scopulorum Engelm.) forests of the American Southwest and provides critical food and habitat for many wildlife species. During the last century, this stratum has been affected by livestock grazing and by increased dominance of overstory trees. We sampled a unique grazing exclosure to examine the relative importance of long-term livestock grazing (grazed or ungrazed) and habitat (park or tree) on the understory community. We sampled 3 plots of 192 contiguous quadrats (each quadrat 0.5 m 2 ) in each of the 4 treatment combinations, for a total of 2304 quadrats. Species-area curves were generated by aggregating quadrats into nonoverlapping areas at grain sizes of 0.5 to 576 m 2 . The effects of habitat and grazing on species density were evident at very different scales. Species density was higher in park than tree plots at scales ≤32 m 2 but did not differ between habitats at larger scales. Species density differed minimally between grazed and ungrazed treatments at small grains, but grazed plots contained more species than ungrazed plots at larger grains. Grazing treatments differed at smaller grains (to 4–8 m 2 ) than did habitats (to 32 m 2 ), with respect to density of native species and graminoids. Grazed plots had more exotic species than ungrazed plots at all grain sizes, though few exotics were present. Twenty-two species were identified as indicator species associated with habitats and/or grazing treatments. Evaluations of plant community response to treatments would be improved by accounting for the grain at which data have been collected and analyzed and by identifying indicator species associated with various treatments. These data would enable more-informed conservation and management decisions.     

Effect of Timing of Grazing on Soil-Surface Crytogamic Communities in a Great Basin Low-Shrub Desert: A Preliminary Report

Cover and species richness of vascular and cryptogamic components of the plant community were inventoried in experimental grazing paddocks at the USDA/FS Desert Range Experimental Station, Millard County, Utah. The grazing treatments considered have been applied continuously for over 50 years. The effects of heavy (ca 17 sheep days/acre) grazing treatment applied in two different seasons (early winter versus a split between early and late winter) differed significantly between seasons. Cryptogamic cover and cryptogamic species richness both showed larger decreases under early-late as opposed to early winter only grazing. Vascular plant cover (relative to controls) was also reduced by early-late winter grazing, but not to a significant degree. Late season grazing, likewise, had no significant effect on number of vascular species per transect.     

Effects of rotational grazing on rodents and raptors in a coastal grassland

We conducted a 4-year experiment to assess the impacts of rotational cattle grazing on rodents and raptors in a mesic coastal grassland in northwestern California. Live-trapping indicated that rodent abundance declined by 69% on the grazed area and increased by 14% on the ungrazed area. Raptor use of the grazed area declined by 15% and increased on the ungrazed area by 63%. Measures of giving-up density indicated that rodents perceived a 25% higher predation risk on grazed area than on ungrazed area, but raptor hunting surveys indicated that risk of depredation from raptors was 2.5 times lower in the grazed habitat, suggesting that rodents use indirect vegetative cues to assess risk.     

Recovery rates of cryptobiotic crusts: inoculant use and assessment methods

Recovery rates of cyanobacterial-lichen soil crusts from disturbance were examined. Plots were either undisturbed or scalped, and scalped plots were either inoculated with surrounding biological crust material or left to recover naturally. Natural recovery rates were found to be very slow. Inoculation significantly hastened recovery for the cyanobacterial/green algal component, lichen cover, lichen species richness, and moss cover. Even with inoculation, however, lichen and moss recovery was minimal. Traditional techniques of assessing recovery visually were found to underestimate time for total recovery. Other techniques, such as extraction of chlorophyll a from surface soil and measurement of sheath material accumulation, were used and are discussed.     

Cryptogams, vascular plants, and soil hydrological relations: some preliminary results from the semiarid woodlands of eastern Australia

Grazed and ungrazed sites were examined in a semiarid woodland in eastern Australia to determine relationships within various types of cryptogams, and the role of cryptogams in pasture dynamics, infiltration, and water erosion. Strong relationships were found between vascular plant cover and cover of cryptogams for nine rangeland sites over an 18-month period. In the absence of vascular plants, sites with low cover of cryptogams were dominated by algae. The presence of a cryptogamic crust had no significant effect on infiltration at ungrazed sites but significantly increased infiltration at some grazed sites. Splash erosion was very low on soils with at least 50% cryptogam cover. Below this level splash erosion increased markedly, along with the proportion of fine sediments lost.     

Effects of cattle grazing on North American arid ecosystems: a quantitative review

A quantitative review was conducted of the effects of cattle grazing in arid systems on 16 response variables ranging from soil bulk density to total vegetative cover to rodent species diversity. Various studies from North American arid environments that used similar measures for assessing grazing effects on the same response variables were used for the review; each study was assigned to serve as a single data point in paired comparisons of grazed versus ungrazed sites. All analyses tested the 1-tailed null hypothesis that grazing has no effect on the measured variable. Eleven of 16 analyses (69%) revealed significant detrimental effects of cattle grazing, suggesting that cattle can have a negative impact on North American xeric ecosystems. Soil-related variables were most negatively impacted by grazing (3 of 4 categories tested were significantly impacted), followed by litter cover and biomass (2 of 2 categories tested), and rodent diversity and richness (2 of 2 categories tested). Vegetative variables showed more variability in terms of quantifiable grazing effects, with 4 of 8 categories testing significantly. Overall, these findings could shed light on which suites of variables may be effectively used by land managers to measure ecosystem integrity and rangeland health in grazed systems.     

Site characteristics and habitat requirements of the endangered dwarf bear-claw poppy ( Arctomecon humilis Coville, Papaveraceae)

Arctomecon humilis Cov. is a narrow endemic, confined to gypsiferous substrates derived from the Shnabkaib Member of the Moenkopi Formation in southwestern Utah. The characteristics of seven A. humilis sites were studied to define habitat requirements of the species. Both physical and biotic aspects of the habitat were analyzed; geology, soil chemistry, and physical properties, as well as vascular and nonvascular plant communities, were studied. Chemical and physical properties of these soils vary considerably from those supporting adjacent desert-shrub communities. The dominant vascular species on A. humilis sites are shadscale and Mormon tea. It was found that A. humilis occurred in portions of the habitat where shrubs were relatively less dense. A soil-surface, cryptogamic community contributed 84% or more of the total living cover on sites that supported A. humilis . Species composition of the cryptogamic community was highly similar among sites. Likewise, composition of the cryptogamic cover was similar when random samples were compared with samples centered on A. humilis plants. The Purgatory Flat site, which does not support the poppy, seems inseparable from A. humilis sites, in respect to soil characteristics and composition of the associated plant cover. There is reason to believe an A. humilis population could be established at the Purgatory Flat site.     

Species richness and California voles in an annual and a perennial grassland

Populations of a common burrowing rodent, Microtus californicus (the California vole), thrive in ungrazed or lightly grazed grasslands in coastal California. Two sites ungrazed by livestock, one dominated by native perennial grasses and another dominated by invasive annuals, were evaluated over 2 consecutive years for the relationship between plant species richness and location of M. californicus burrow entrances (burrows). Plant species and burrows were sampled as present or absent in contiguous 1-m 2 quadrats on a 100-m 2 grid. Quadrats with burrows averaged significantly more plant species than quadrats without them (11.3 vs. 9.9 species, P < 0.001). Burrows found in 1996 were not correlated with species richness in 1995, suggesting that voles affect richness rather than seek it out. Vole burrow locations showed significant clumping on the annual site and trended toward clumping on the perennial site in both 1995 and 1996. Because voles seem to create a clumped pattern with their burrow entrances, the associated increase in plant species richness may have a strong effect on the overall structure of the plant community. A quantitative comparison of the 2 sites showed that the plant matrix of the perennial site contained flora of the annual site. This similarity in plant species composition may allow for similar treatment of our 2 types of sites and potentially other California grasslands. Undetected increases in vole populations with livestock grazing reduction may account for the erratic results from grasslands management research and the inconsistent success of derived management practices.     

Effects of livestock grazing exlosure on aquatic macroinvertebrates in a montane stream, New Mexico

Aquatic macroinvertebrate populations inhabiting reaches of a stream within areas excluded from livestock grazing for a decade were markedly different from those in grazed areas when density, biomass, biotic condition indices, and mean chi square indices of the two populations were compared. Increased densities and biomasses of more tolerant forms of macroinvertebrates were observed in grazed reaches. Because pretreatment data were not available, differences in macroinvertebrate populations and relative tolerances of taxa in grazed and ungrazed areas could be as easily attributed to linear changes in stream habitat as to removal of domestic livestock. Results of this study have implications for the design of future research on the effects of livestock grazing on stream environments and biota: (1) baseline/pretreatment information is prerequisite, and (2) the study should take a watershed (ecosystem) approach.     

Lichens of soil crust communities in the Intermountain Area of the western United States

Lichens are common components of microbiotic soil crusts. A total of 34 species from 17 genera are reported from soil crust communities throughout the Intermountain Area. Distribution of terricolous lichens is determined by various physical and biological factors: physical and chemical characteristics of the soil, moisture regimes, temperature, insolation, and development and composition of the vascular plant community. Some species demonstrate a broad ecological amplitude while others have a more restricted distribution. All growth forms are represented; however, the vast majority of soil crust lichens are squamulose (minutely foliose). Fruticose species are least abundant. In exposed, middle-elevation sites vagrant (detached) species are common. This paper describes and discusses terricolous lichen communities of desert habitats of the intermountain western United States. Effects of various human-related activities including grazing, wildfire, air pollution, and recreation vehicles on soil crust lichens are discussed. Gypsoplaca macrophylla (Zahlbr.) Timdal, a rare squamulose lichen which occurs on gypsifersous soils, was recently collected in Emery County, Utah, and is reported as new to the state.     

Comparative habitat and community relationships of Atriplex confertifolia and Sarcobatus vermiculatus in central Utah

Thirty-four study sites were established in shadscale ( Atriplex confertifolia [Torr. & Frem.] Wats.) and greasewood ( Sarcobatus vermiculatus [Hoov. Torr. in Emory) communities bordering Utah Lake in central Utah. Differences in species composition, vegetation, and soil characteristics were assessed. Significant differences in soil factors between the two communities were found for sand, calcium, manganese, zinc, and copper. Soluble salts and sodium concentrations were generally higher in the greasewood type, but differences were not significant. Major differences were found in understory species, with burr buttercup ( Ranunculus testiculatus Grantz) showing significantly greater cover in the shadscale community and cheatgrass ( Bromus tectorum L.) showing significantly greater cover in the greasewood community.     

Foliage biomass and cover relationships between tree- and shrub-dominated communities in pinyon-juniper woodlands

Woodlands dominated by singleleaf pinyon ( Pinus monophylla Torr.and Frem.) and Utah ( Juniperus osteosperma [Torr.] Little) cover extensive areas in the Great Basin and Southwest. Both species are aggressive and can nearly eliminate the previous shrub-dominated community. Successional pathways from shrub-dominated communities before tree establishment to the tree-dominated communities that follow are known only for a few specific sites. How site growing conditions affect successional patterns needs further study. We compared the relationship of foliage biomass and percentage of cover between paired shrub-dominated and tree-dominated plots over several sites. Sites studied are from different elevation and topographic conditions on one mountain range. Foliage biomass in shrub-dominated plots had about a three-to-one variation over the range of site conditions sampled. Tree-dominated plots varied by about two-to-one. Cover in shrub-dominated plots had a four-to-one variation; cover in the tree-dominated plots varied by about two-to-one. Total foliage biomass in both tree- and shrub-dominated plots correlated best with the site index of height at 200 years of age. Variation in percentage of cover in both tree and shrub-dominated plots correlated best with elevation. Foliage biomass variation in shrub-dominated plots was proportional to the variation in the paired tree-dominated plots. A similar proportional relationship was present for percentage of cover between paired tree- and shrub-dominated plots. Foliage biomass was more sensitive to topographic differences than to cover. Variation in plant species sampled in the shrub-dominated plots correlated with total foliage biomass of the same plots. Species sampled also correlated with pinyon height at 200 years of age and total foliage biomass in the paired tree-dominated plots.