Spatial patterns of plant communities and differential weathering in Navajo National Monument, Arizona

Vegetation patterns in Navajo National Monument, Arizona, were studied over a five-year period from 1977 to 1981. Twelve distinct plant community types occur within the boundaries of the park. These communities are characterized and the dominant plant species of each are recorded. The relationships of parent material, soils, and moisture to plant communities are also discussed. It appears that discrete communities occupy soils of different characteristics, particularly with respect to amount of weathering of parent material.         

Plants of Navajo National Monument

The floristic composition of Navajo National Monument is presented. The flora of the monument consists of 293 species of vascular plants, representing 177 genera and 66 families. The species are scattered throughout 12 plant communities found within the monument. The flora is dominated by forbs (60 percent), followed by shrubs (16 percent), grasses (12 percent) and trees (5 percent). Betatakin canyon exhibits the greatest floristic richness with 223 species being represented and 123 species found only in that area. This is due primarily to two factors: (1) the greater variety of habitats available in the area, and (2) the much longer time the area has been protected from grazing and other manmade disturbance.     

Influence of cryptogamic crusts on moisture relationships of soils in Navajo National Monument, Arizona

Normal 0 false false false EN-US X-NONE X-NONE MicrosoftInternetExplorer4 st1\:*{behavior:url(#ieooui) } /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Table Normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-priority:99; mso-style-qformat:yes; mso-style-parent:""; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:11.0pt; font-family:"Calibri","sans-serif"; mso-ascii-font-family:Calibri; mso-ascii-theme-font:minor-latin; mso-fareast-font-family:"Times New Roman"; mso-fareast-theme-font:minor-fareast; mso-hansi-font-family:Calibri; mso-hansi-theme-font:minor-latin; mso-bidi-font-family:"Times New Roman"; mso-bidi-theme-font:minor-bidi;} Cryptogamic soil crusts of Betatakin Canyon in Navajo National Monument were investigated to understand the influence of such crusts on soil moisture relationships and potential sediment production. Crusts sampled were part of the pinyon-juniper community and were studied in paired units. The presence of crusts on soils significantly increased the depth of water penetration and decreased runoff. Soils showed reduced infiltration of water where lichen and algal crusts were present and enhanced infiltration rates where mosses were present. Crusts appear to cause surface sealing and therefore likely reduce surface evaporation rates as well.       

Bird community composition in relation to habitat and season in Betatakin Canyon, Navajo National Monument, Arizona

Bird species density, diversity, and species richness in relation to habitat and seasonal variations were studied in the Betatakin Canyon area of Navajo National Monument, Arizona. The two most prominent habitat types are a riparian forest deep in the canyon bottom and a mature pinyon-juniper woodland on the mesas and slopes above the canyon. One hundred thirty-five species of birds were encountered during the study and recorded by season and habitat. The avifauna assemblages demonstrate definite habitat selection into groups associated with the riparian and pinyon-juniper woodland communities. Diversity, density, and species richness were greatest in the riparian habitat during the spring and summer months. The differences in bird community composition were greater between seasons than between habitats. Pinyon pine and Gambel oak were highly selected as perch sites in the pinyon-juniper and riparian areas, respectively.     

Hosts of juniper mistletoe at Walnut Canyon National Monument, Arizona

Four species of junipers, Juniperus osteosperma, J. scopulorum, J. monosperma, and J. deppeana, occur in mixed stands at Walnut Canyon National Monument, Arizona. All are parasitized by Phoradendron juniperinum, but the mistletoe was most common on J. osteosperma and least common on J. scopulorum. Fernbush ( Chamaebatiaria millefolium, Rosaceae) was a very rare host.     

Grass spider microhabitat use in Organ Pipe Cactus National Monument, Arizona

The grass spider ( Agelena naevia ), commonly found in Organ Pipe Cactus National Monument, Arizona, uses rodent burrows located under a shrub canopy more frequently for web construction than burrows located in the open. The average number of prey available in canopy microhabitat was greater than in open microhabitat, and unequal prey abundance may explain spider microhabitat use.        

Birds of northern Black Mesa, Navajo County, Arizona

Two hundred forty-one species of birds have been identified from northern Black Mesa, Arizona. This region's avifauna was poorly known until the late 1970s when large-scale coal mining began. Vegetation of the region is predominantly Great Basin desert scrub, pinyon-juniper woodland, and mixed-conifer woodland. The latter vegetation supports an assemblage of isolated montane bird species unique to the region. Numerous environmental changes have recently affected the bird life of Black Mesa. These include large-scale type conversions (pinyon-juniper clearing and surface mining), pond construction, and establishment of exotic vegetation.     

Diatoms of Oregon Caves National Monument, Oregon

The diatom flora of the Oregon Caves National Monument, Josephine Co., Oregon, was investigated. Diatoms were distributed throughout the cave system with abundance and species diversity depending upon moisture, light, availability of mineral nutrients, and proximity to cave openings. Twenty-six taxa were identified and described.     

Terrestrial vertabrates of Scotts Bluff National Monument, Nebraska

A survey of terrestrial vertebrates was conducted at the Scotts Bluff National Monument (SBNM), Nebraska, to determine species composition, relative abundance, and distribution by habitat of the existing amphibians, reptiles, birds, and mammals. Various sampling methods were used to detect animal species richness and to estimate population densities. The seven major habitat types at SBNM contained 4 species of amphibians, 8 reptiles, 96 birds, and 28 mammals. The only endangered or threatened species observed was a dead bald eagle ( Haliaeetus leucocephalus ). The racer ( Coluber constrictor ) and prairie rattlesnake ( Crotalus viridis ) were the two most abundant snakes. The prairie falcon ( Falco mexicanus ) was a common raptor that nested in Scotts Bluff. Sympatric populations of mule deer ( Odocoileus hemionus ) and white-tailed deer ( Odocoileus virginianus ) appeared to be hybridizing. The riverine woodland contained the greatest number of fauna species with only 4% surface area of SBNM. Sharp-tailed grouse ( Tympanuchus phasianellus ) and pronghorn antelope ( Antilocapra americana ), absent from the Monument, are two potential native species for reintroduction.     

Preliminary mammal survey of Fossil Butte National Monument, Wyoming

The mammalian community of Fossil Butte National Monument, Wyoming, consisted of at least 45 species. Snap trap sampling of the smaller species combined with sight and sign observations of the larger species formed the basis of this preliminary survey in the summers of 1974 and 1975. Snap - trapping samples (2880 trap days) yielded 44 least chipmunks and 89 deer mice. The distribution of all species was characterized by plant community occupancy. Heavy domestic livestock use of the communities nearest water has resulted in severe alteration in community structure; this has probably affected the distribution and abundance of at least some of the smaller mammals.     

Vegetation response to prescribed fire in Dinosaur National Monument

Much of western North America is dominated by dense, monotypic, late seral stands of big sagebrush ( Artemisia tridentata Nutt.). These stands often have depauperate understories with limited species richness, diversity, and herbaceous cover. The National Park Service at Dinosaur National Monument, Colorado, is using both strategic and natural prescribed fire in Wyoming big sagebrush ( Artemisia tridentata ssp. wyomingensis Beetle and Young) communities to foster intra-community (α -scale) and landscape diversity. This study analyzed an accumulated foliar cover data set between paired burn and control areas on 6 different sites during the last 20 years. Across the monitoring period, mean total vegetation cover of all combined sites was 44% control and 42% burn. Total vegetation cover in burn areas was higher than or equal to paired control areas within 2-3 years post-burn. Shrubs were essentially eliminated in burn areas, but perennial grass cover was 10-35% higher. Mean number of species on all sites and years combined was 17 control and 18 burn. Species richness was different on only 1 site-year, Dry Woman 1995 ( P = 0.001, 15 control, 9 burn). Species similarity by site and between treatments ranged from 44% to 75%. Differences in Shannon-Weiner diversity index values between paired sites occurred in 6 of 20 years ( P < 0.05). Index value differences on these 6 sites were due to a large annual grass component in burn areas. Prescribed burning successfully shifted late successional sagebrushdominated communities to earlier herbaceous-dominated successional stages without lowering total vegetation cover, while maintaining -scale diversity and species richness.     

Riffle beetles (Coleoptera: Elmidae) of Death Valley National Monument, California

Three species of Elmidae occur in Death Valley National Monument: Stenelmis calida is in three springs in the Ash Meadows area; Microcylloepus formicvideus is only in Travertine Springs; and Microcylloepus similis is in several springs throughout Death Valley and Ash Meadows. Only permanent springs support elmids. Considerable morphological variation occurs in the disjunct populations of M. similis . The evolution of elmids in Death Valley National Monument is equivalent to that of the local pupfish ( Cyprinodon spp.).     

Fire effects on small mammal communities in Dinosaur National Monument

Resource specialists at Dinosaur National Monument utilize both planned and unplanned wildland ignitions in big sagebrush ( Artemisia tridentata )--dominated plant communities to restore successional processes, maintain vegetation vigor, and promote diversified landscapes. Short- and long-term effects of prescribed burning on small mammal populations are not well understood. The objectives of this study were to (1) compare small mammal species richness, similarity, and diversity between paired burned and unburned treatment plots, and (2) assess long-term trends of small mammal community responses to burning. Five paired burned/unburned sites having similar vegetation, soils, elevation, and annual precipitation were selected. Prescribed or natural fires occurred on all sites between 1981 and 1995, followed by periodic small mammal removal (trapping) sampling (1--12 years post-burn). Small mammal species richness and diversity were generally higher on unburned than burned plots across post-burn sampling years and sites. Increased abundance of the deer mouse ( Peromyscus maniculatus ) following burning resulted in decreased community diversity. Species similarities between burned and unburned plots were low in early post-burn years but increased in later post-burn years. Short-term shifts in diversity and species similarity resulted from increased deer mouse abundance after burning. However, burning had no long-term impact on species richness and similarity, indicating minimal effect to other small mammal species.     

Black-tailed prairie dog population dynamics at Scotts Bluff National Monument, Nebraska: a 28-year record

Black-tailed prairie dogs, Cynomys ludovicianus, now inhabit a small fraction of their original range in the Great Plains. We monitored a population of black-tailed prairie dogs at Scotts Bluff National Monument, Nebraska, from colonization in 1981 until 2009 (28 years). Colony boundaries were mapped by delineating clip lines and active burrows; population densities were estimated via visual counts. Estimates of total population size revealed 4 distinct periods of changing dynamics: (1) a linear increase, (2) a decline and prolonged depression, (3) an exponential increase, and (4) a period of high variability. Area occupied revealed similar, although less-defined trends, whereas densities fluctuated greatly (8&ndash;80 individuals &middot; ha &ndash;1 ). Even after almost 30 years, this population remains relatively small. Decreases in the population may have been due, in part, to predation by badgers, although sylvatic plague cannot be ruled out. Black-tailed prairie dogs are recognized as keystone grassland species, and attempts are underway to reintroduce them to parts of their historic range. Our data suggest that black-tailed prairie dogs possess high potential for rapid population growth and decline, regardless of colony size. Therefore, either human-assisted or natural dispersal events may be important in establishing colonies in suitable habitat. El perrito de pradera de cola negra, Cynomys ludovicianus, habita en una peque&ntilde;a fracci&oacute;n de su distribuci&oacute;n original en las Grandes Llanuras. Monitoreamos una poblaci&oacute;n de perritos de pradera de cola negra en Scotts Bluff National Monument, Nebraska, por 28 a&ntilde;os, desde la colonizaci&oacute;n en 1981 hasta 2009. Trazamos los l&iacute;mites de las colonias usando de referencia las l&iacute;neas de ramoneo y las madrigueras activas, y estimamos la densidad de las poblaciones mediante un recuento visual. Los estimados de poblaci&oacute;n total revelaron cuatro per&iacute;odos distintos de din&aacute;micas cambiantes: (1) un aumento lineal, (2) una disminuci&oacute;n y depresi&oacute;n prolongada, (3) un aumento exponencial y (4) un per&iacute;odo de alta variabilidad. El &aacute;rea ocupada revel&oacute; tendencias similares, aunque menos definidas, mientras que las densidades fluctuaron mucho (8&ndash;80 individuos &middot; ha &ndash;1 ). A&uacute;n despu&eacute;s de casi 30 a&ntilde;os, esta poblaci&oacute;n permanece relativamente peque&ntilde;a. Los decrementos en la poblaci&oacute;n pueden haber sido provocadas en parte por la depredaci&oacute;n por tejones, aunque no se puede descartar como causa la plaga silv&aacute;tica. El perrito de pradera de cola negra se considera una especie clave de la pradera, y est&aacute;n en proceso esfuerzos por reintroducirlos en partes de su distribuci&oacute;n hist&oacute;rica. Nuestros datos indican que los perritos de pradera de cola negra tienen alto potencial de crecimiento y disminuci&oacute;n de su poblaci&oacute;n, sin importar el tama&ntilde;o de la colonia. Por lo tanto, los eventos de dispersi&oacute;n, ya sean naturales o con asistencia humana, podr&iacute;an ser importantes para establecer colonias en h&aacute;bitats adecuados.     

Estimates of the incidence of mistletoes in pinyon-juniper woodlands of the Coconino National Forest, Arizona

The relative incidence of mistletoes found in pinyon-juniper woodlands is estimated for the Coconino National Forest, Arizona, using a roadside survey. Approximately 50% of the pinyon-juniper woodlands surveyed were infested with juniper mistletoes ( Phoradendron juniperinum or P. capitellatum ). In contrast, only about 12% of the areas surveyed were infested with pinyon dwarf mistletoe ( Arceuthobium divaricatum ). In these infested woodlands, 70% with juniper mistletoes and 44% with pinyon dwarf mistletoe were lightly infested. Most areas where pinyon dwarf mistletoe populations were observed (90%) also had juniper mistletoe ( P. juniperinum ) present, but many juniper mistletoe populations observed had no pinyon dwarf mistletoe (50%), although pinyon pine was common at those locations. Therefore, pinyon dwarf mistletoe is frequently co-distributed with juniper mistletoe as has been suggested by other investigators, but juniper mistletoe frequently occurs where there is no pinyon dwarf mistletoe present.     

Microbial spatial heterogeneity in microbiotic crusts in Colorado National Monument. II Bacteria

Microbial spatial heterogeneity and bacterial composition of homogeneous surface soils showing evident microbiotic crust development located under Utah juniper canopies in Colorado National Monument were studied. Four distinct homogeneous sampling sites positioned along a 12-m transect were compared on the basis of bacterial density and taxa composition. Bacterial densities showed a range differing by several orders of magnitude. In comparisons of adjacent soil samples or samples taken from two different sampling sites within close proximity, adjacent samples showed no more similarity to each other on the basis of bacterial density or taxa composition than did more widely separated samples. The bacterial composition of these soils included members of the genera Bacillus , Micrococcus , and Arthrobacter . Actinomycetes were also prevalent.     

Fishtail Mesa: a vegetation resurvey of relict area in Grand Canyon National Park, Arizona

Relict sites are geographically isolated areas that are undisturbed by direct and indirect human influences. These sites facilitate long-term ecological monitoring by providing a reference for gauging impacts occurring elsewhere. Knowledge gained through comparing vegetation change on matched relict and proximal disturbed areas can help partition the causes of change into natural and human-produced components. Fishtail Mesa in Grand Canyon National Park is a 439-ha relict site that is inaccessible to domestic livestock. Human visitation is infrequent and irregular, and fires have never been suppressed or managed. In 1958, U.S. Forest Service range scientists conducted a survey of Fishtail Mesa to gather reference data on vegetation, wildlife, and soils. Vegetation sampling was conducted using a method called the ";";elb.";"; We returned to Fishtail Mesa in May 1996 to perform a general vegetation and floristic survey, assess the extent of vegetation change after 38 years, and evaluate the suitability of the site as a location for long-term surveillance of ecological change. Fishtail Mesas vegetation consists primarily of a Pinus edulis (pinyon) and Juniperus osteosperma (Utah juniper) woodland with an Artemisia tridentata (sagebrush) understory, or tree-type (310.9 ha), and an Artemisia and Poa fendleriana (mutton grass) steppe, or shrub-type (127.5 ha). Since 1958 vegetation changes in both shrub- and tree-types have been limited to only a few species. In the shrub-type we detected slight increases from 1958 to 1996 in both Pinus and Juniperus , and reexamination of 1958 photo sites confirmed that Pinus and Juniperus are reoccupying the shrub-type. Artemisia cover declined from 1958 to 1996, whereas Poa increased from near trace amounts in 1958 to moderate cover in 1996. In the tree-type, Poa has increased from 1958 to 1996, while Artemisia , Juniperus , and Pinus showed no apparent change. Other species such as Ephedra torreyana (Torrey joint-fir), Opuntia polyacantha (prickly pear), and Gutierrezia sarothrae (snakeweed) have decreased. Vegetation analysis aided by TWINSPAN revealed that the shrub-type is defined more on the basis of absence of Pinus and Juniperus rather than any special association of differential species with a high preference for this type. We interpret the ";";invasion";"; of the shrub-type by Pinus and Juniperus as a ";";reoccupation.";"; Indirect ordination using DECORANA inferred 2 environmental gradients, a moisture gradient and perhaps a substrate texture gradient, that appeared to influence vegetation distribution on Fishtail Mesa. Fishtail Mesa is a valuable relict area for studying the effects of livestock grazing and prescribed fire. It should be designated a Federal Research Natural Area based on its vegetation communities, size, and protection afforded by its location in Grand Canyon National Park.     

Microbial spatial heterogeneity in microbiotic crusts in Colorado National Monument. I. Algae

Spatial heterogeneity in visually similar sites under Utah juniper canopy in Colorado National Monument was examined. Sample sites were arranged in a transect 24 m long, such that distances between samples were 0.013 m, 0.03 m, 1.0 m, 12 m, and 24 m. Twenty-five taxa of algae were observed, mostly belonging to Cyanophyta. Algal density varied by more than an order of magnitude within the 46 samples examined. The coefficients of variation for each distance class were very similar, demonstrating that algal patchiness can be as significant on a scale less than 0.013 m as it is on a scale of 24 m. Goodall's random pairing analysis of spatial pattern supported this conclusion by indicating that the minimal area for sampling soil algal crust populations at this site was equal to or less than 0.013 m. Because of the microscale heterogeneity in algal communities in this study, we recommend that future researchers take composite samples if they wish to quantify algae of microbiotic crusts.     

Vegetation response to fire and postburn seeding treatments in juniper woodlands of the Grand Staircase-Escalante National Monument, Utah

We compared 3 naturally ignited burns with unburned sites in the Grand Staircase-Escalante National Monument. Each burn site was restored with native and nonnative seed mixes, restored with native seeds only, or regenerated naturally. In general, burned sites had significantly lower native species richness (1.8 vs. 2.9 species), native species cover (11% vs. 22.5%), and soil crust cover (4.1% vs. 15%) than unburned sites. Most burned plots, seeded or not, had significantly higher average nonnative species richness and cover and lower average native species richness and cover than unburned sites. Regression tree analyses suggest site variation was equally important to rehabilitation results as seeding treatments. Low native species richness and cover, high soil C, and low cover of biological soil crusts may facilitate increased nonnative species richness and cover. Our study also found that unburned sites in the region had equally high cover of nonnative species compared with the rest of the Monument. Cheatgrass ( Bromus tectorum ) dominated both burned and unburned sites. Despite the invasion of cheatgrass, unburned sites still maintain higher native species richness; however, the high cover of cheatgrass may increase fire frequency, further reduce native species richness and cover, and ultimately change vegetation composition in juniper woodlands.     

European earwig, Forficula auricularia L. (Dermaptera: Forficulidae), at the Hanford Reach National Monument, Washington State

The European earwig, Forficula auricularia L., was surveyed using pitfall traps at 3 sites at the Hanford Reach National Monument in south central Washington State. Pitfall traps were collected weekly from April 2002 through April 2003. The earwig was consistently taken during all months of the year at a disturbed, weedy site along the Columbia River (White Bluffs Ferry site) but was rare or not collected in 2 less-disturbed shrubsteppe habitats. Highest numbers occurred during April&ndash;May, when immatures accounted for the majority of the catch; immatures reached the adult stage during mid-July, and the species is univoltine at the site. Possible reasons why earwigs have not colonized the monument&rsquo;s shrubsteppe habitat include the arid climate with lack of available moisture, especially for breeding purposes, and a lack of burrow sites. La tijereta Europea, Forficula auricularia L., fue estudiada usando trampas de ca&iacute;da en tres sitios del Hanford Reach National Monument en el sur del estado de Washington. Se colectaron los contenidos de las trampas de ca&iacute;da semanalmente desde abril de 2002 hasta abril de 2003. La tijereta apareci&oacute; constantemente durante todos los meses del a&ntilde;o en un sitio perturbado de vegetaci&oacute;n herb&aacute;cea en la orilla del R&iacute;o Columbia (el lugar del antiguo transbordador White Bluffs) pero su colecta fue escasa o nula en dos h&aacute;bitats de estepa arbustiva menos perturbados. Los mayores n&uacute;meros ocurrieron durante abril y mayo cuando los organismos inmaduros representaron la mayor&iacute;a de los espec&iacute;menes capturados; &eacute;stos alcanzaron la etapa adulta a mediados de julio y la especie es univoltina en este sitio. Posibles causas de que las tijeretas no hayan colonizado el h&aacute;bitat de estepa arbustiva del Monumento podr&iacute;an incluir su clima &aacute;rido carente de agua disponible, especialmente para la reproducci&oacute;n, y una falta de sitios para madrigueras.