Colorado's rare flora

Distribution, status, habitat, elevational range, original literature citations, synonymy, and specimens deposited at Brigham Young University (BRY), University of Colorado (COLO), Colorado State University (CS),Denver Botanical Garden (KHD), Kansas State University (KSC), University of Kansas (KANU), Missouri Botanical Garden (MO), and the Rocky Mountain Herbarium (RM) are given for 79 species of rare Colorado plants. Species federally listed by the U.S. Fish and Wildlife Service under the Endangered Species Act, candidates for listing, and species that qualify as federal candidates because of limited range, small populations, or known threats are included. Suggested changes in status are offered where appropriate. Maps and tables indicating the Colorado range of these species are provided.     

Checklist of the Odonata of Colorado

One hundred and one species of Odonata are recorded for Colorado, including 58 species of Anisoptera (of which 4 are new state records) and 43 Zygoptera (9 new state records). County records are indicated for each. Three major elements are recognized; predominantly eastern species occurring in the plains and mountain valleys east of the Rocky Mountains, Canadian Zone elements occurring principally above 2,440 m (8,000 ft) in the mountains, and Great Basin and western species occurring chiefly west of the mountains. The list is based primarily on the collections at Colorado State University and the University of Colorado, with additional records from the literature and from other collections.     

Colorado ground beetles (Coleoptera: Carabidae) from the Rotger Collection, University of Colorado Museum

Ground Beetles from Rotger's collection of Colorado specimens have been identified, principally by the author, and a faunal list of 161 species from 80 localities is presented. The list includes 35 species not previously recorded from Colorado. Comparisons are made with Armin's (1963) carabid list from Boulder County and the diversity of species along transects through four elevational zones from the plains to the alpine.     

On the Distribution of Utah's Hanging Gardens

This is a summary monograph of the hanging gardens as they occur in the Colorado River and Virgin River portion of the Colorado Plateau in Utah. Discussed in this paper are the hanging gardens, their geography, geomorphology, aspects of distribution and diversity, and principal vascular and algal plant species. Animal trapping studies and pland productivity aspects are reviewed.     

Astragalus (Leguminosae): nomenclatural proposals and new taxa

This is a summary monograph of the hanging gardens as they occur in the Colorado River and Virgin River portion of the Colorado Plateau in Utah. Discussed in this paper are the hanging gardens, their geography, geomorphology, aspects of distribution and diversity, and principal vascular and algal plant species. Animal trapping studies and pland productivity aspects are reviewed.     

Caddisfly (Trichoptera) assemblages along major river drainages in Arizona

Seventy-two caddisfly species in 36 genera and 15 families were collected along the Little Colorado, Verde, Gila, and Colorado rivers in Arizona. Brachycentrids, hydropsychids, limnephilids, and uenoids made up nearly 50% of the assemblage in forested sites above 2200 m. Hydropsychids, hydroptilids, and leptocerids made up 75% of the assemblage in grassland watersheds between 2200 and 1000 m, and 80% at sites below 1000 m in desert watersheds. Species richness averaged 16 at sites above 2200 m elevation, 7 between 2200 and 1000 m, and 3 below 1000 m. H' diversity was typically >3 at sites above 2200 m and <2.5 below 2200 m. Each river had distinct faunas likely due to isolation of drainages across semiarid landscapes. Low S&oslash;rensen similarity values were measured for caddisfly assemblages in the highly regulated Colorado River and the other 3 rivers. Only 17 species were collected at 14 sites along 700 km of the Colorado River. Indicator caddisfly species are provided for altered and less disturbed drainages. We added 7 species not previously reported in Arizona for a total of 154 caddisfly species in the state. We provide a baseline of caddisfly indicators for monitoring changes in ecosystem health during the predicted long-term drought and population expansion in southwestern USA.     

Archaeological record of native fishes of the lower Colorado River: how to identify their remains

Archaeological sites in the Salton Basin of southeastern California and along the lower Colorado River provided opportunities to determine which fish species were present prior to extirpations, environmental degradation, and the recession of Lake Cahuilla. These remains also represent the fishes exploited by Native Americans. Bonytail ( Gila elegans ), razorback sucker ( Xyrauchen texanus ), Colorado pikeminnow ( Ptychocheilus lucius ), striped mullet ( Mugil cephalus ), and machete ( Elops affinis ) have been recovered from 117 sites in the Salton Basin, once filled by the Colorado River forming Lake Cahuilla. Bonytail and razorback sucker comprise nearly 99% of the remains. Along the lower Colorado River itself, fragmentary elements of bonytail, razorback sucker, Colorado pikeminnow, and roundtail chub ( G. robusta ) have been recovered, documenting a disappearing native fish fauna. Anatomical details are described that permit identification of diagnostic materials commonly recovered during archaeological excavations.     

Reestablishment of Stenogonum Nutt. (Polygonaceae)

The genus Stenogonum Nutt. (Polygonaceae) is reestablished and distinguished from Eriogonum Michx. with which it has been synonymized since 1853. The genus, as outlined here, differs from Eriogonum in having two whorls of three foliaceous bracts surrounding the flowers instead of the normal, united tubular involucre. Stenogonum contains two species, S. flexum and S. salsuginosum; they are restricted primarily to the Colorado Plateau of Wyoming southward through eastern Utah and western Colorado to northwestern New Mexico and adjacent northern Arizona. Both species are illustrated and their respective ranges mapped.     

New records and range extensions of species of Dipogon (Hymenoptera, Pompilidae) in Colorado

Four species of Dipogon are recorded from Colorado for the first time. These are hurdi Evans, pulchripennis (Cresson), graenicheri Banks, and brevis (Cresson). All are inhabitants of wooded areas and are believed to nest in cavities in trees. Notes are presented on habitat and prey of these 4 species and of the recently described species kiowa Evans.     

New species and a new combination of Mentzelia section Bartonia (Loasaceae) from the Colorado Plateau

A new species, Mentzelia (sect. Bartonia ) cronquistii , subfamily Mentzelioideae, is described and a new combination, M. (sect. Bartonia ) marginata (Osterhout) is made. These two species of the Colorado Plateau are closely related to each other, but their relationships to other species in section Bartonia are obscure.     

Gizzard shad ( Dorosoma cepedianum ) expansion and reproduction in the Upper Colorado River Basin

Gizzard shad ( Dorosoma cepedianum ) were introduced into the Colorado River Basin circa 1996 via an unintentional contaminated stocking. Gizzard shad were collected throughout much of the Upper Colorado River Basin in 2005, 2006, and 2007. These collections document a rapid range expansion of this introduced species and a threat to intact fisheries.     

New species of oribatids from western Colorado (Acarina: Cryptostigmata, Suctobelbidae)

A new species of Suctobelba, S. tumulata, and a new genus and species, Parisuctobelba septenia, are described from collections of oribatids taken in coal strip-mining spoil banks near Hayden, Colorado. The new species are compared with members of the family from North America and Europe. The new genus is differentiated by a smooth, narrowed sensillum and seven pairs of genital setae. Taxonomic and ecological data are also discussed.     

A new species of Penstemon (Scrophulariaceae) from the Uinta Basin of Utah and Colorado

Named as a new species is Penstemon albifluvis J. L. England. The species is known from the Uinta Basin of Utah and Colorado.     

Incidence of the leech Helobdella stagnalis on the Colorado River in west central Colorado

Fifty-seven fish of three genera were taken from the Colorado River in west central Colorado and examined for external parasites. No external parasites were found on Ictalurus. Seven percent of the Hybognathus and 100 percent of the Catastomus were infected with a Hirudinean of the species Helobdella stagnalis. &nbsp;&nbsp;     

Breeding records for Clark's Grebe in Colorado and Nevada

Described as a new breeding species is Clark's Grebe in Colorado and Nevada.     

Floristic analysis of the southwestern United States

A study was made of the distributions of native, terrestrial, vascular plants occurring in 50 local floras from throughout the Basin and Range and Colorado Plateau physiographic provinces of the southwestern United States. The objectives of the study were to objectively define and describe the floristic elements-assemblages of species with roughly coincident geographic distribution-occurring in the southwestern United States and to determine what such assemblages reveal about the floristic history of the region. The total flora (native, terrestrial species only) of the Southwest is estimated at 5,458 species, 77% of which were recorded in 1 or more of the local floras. Nearly 22% of these species are endemic to the study region. A majority of the species were found to be relatively rare. The average range of a species included only 4 floras, and 90% of the species were recorded from 11 or fewer floras; only 81 species (1.5%) were recorded from 50% or more of the floras. Trees constitute 2% of the regional flora and have the widest average distribution; perennial herbs constitute 59% of the flora and have the most restricted distributions.&nbsp; Factor analysis was used to identify seven floristic elements for the region: a Great Basin element, a Mojavean element, a Colorado Plateau element, a Chihuahuan element, an Apachian element, and a Mogollon element. This factor analysis solution was shown to satisfy criteria of interpretability and consistency. The Mojavean, Colorado Plateau, and Apachian elements are believed to be autochthonous. The other four elements show high overlap in species composition with one or more adjacent regions. Each floristic element is mapped to show its geographic form and distribution. Analysis of these maps shows how the existence of objectively defined floristic elements is not contradictory to either the individualistic view of the distribution of a species or local continuity of vegetation and flora. The rarity of the majority of species and the clear association of floristic elements with rather narrowly circumscribed Holocene environments suggests that many Southwestern species have migrated little and are of rather recent, probably postglacial origin. Geographic &ldquo;principles&rdquo; derived from the distribution patterns of relatively few, widespread, dominant, usually woody species may not be applicable to entire, regional floras. &nbsp;     

First specimen records for Ruffed Grouse in Colorado

We report the first specimen records for Ruffed Grouse ( Bonasa umbellus ) collected in Colorado, provide the location, describe the habitat, and briefly discuss the rarity of this species within the state. Three Ruffed Grouse were collected, an adult male in September 1988 and a juvenile male and adult female in September 1990, on Hoy Mountain, Moffat County, Colorado, within 0.6 km of Utah. The habitat is primarily open Douglas-fir ( Pseudotsuga menziesii ) forest with scattered clumps of quaking aspen ( Populus tremuloides ) and Utah serviceberry ( Amelanchier utahensis ). Habitats to the north, east, and south are primarily dominated by sagebrush ( Artemisia spp.) steppe and pionjuniper ( Pinus spp.-- Juniperus spp.) woodland. We hypothesize that natural expansion of Ruffed Grouse further east, north, and south is prohibited by unsuitable habitat exacerbated by the limited flight range capability of the species.     

Ophiotaenia critica (Cestoda: Proteocephalidae), a parasite of the Colorado River squawfish

A total of 19 cestodes, Ophiotaenia sp., were recovered by deworming from adult Colorado River Squawfish ( Pteichocheilius licius ) maintained at the U.S. National Fish Hatchery at Willow Beach, Arizona. These specimens differ from other described species and are named Ophiotaenia critica.     

Migrating Mormon Crickets, Anabrus simplex (Orthoptera: Tettigoniidae), as food for stream fishes

Migrating bands Mormon crickets ( Ananbrus simplex ) were observed crossing the Green and Yampa rivers in Dinosaur National Monument, Colorado and Utah, in 1986 and 1987. Swimming crickets were swept downstream and eaten by four- endemic and seven introduced fish species. Included were two endangered fishes, Colorado squawfish ( Ptychocheilus lucius ) and humpback chub ( Gila cypha ). Direct and indirect effects to aquatic food webs associated with application of pesticides for Mormon cricket control may pose a threat to these fishes and to man.     

Habitat and plant distributions in hanging gardens of the Narrows, Zion National Park, Utah

Hanging gardens are insular plant communities of the Colorado Plateau. This study examines hanging gardens in the Narrows, Zion National Park, Utah. The floristic similarity of gardens and the presence of species in classes of variables which characterize the habitat are disclosed. Although the gardens are isolated and dissimilar, the individual species are not restricted in the range of habitat found. Floristic dissimilarity cannot be attributed to differences in habitat. These results are compared to studies of hanging gardens in eastern Utah. &nbsp;