Classification of the riparian vegetation of the montane and subalpine zones in western Colorado

A classification of the riparian vegetation of part of western Colorado was developed by cluster analysis and ordination of 115 samples of relatively undisturbed vegetation. The classification of plant associations contains five montane riparian forests, three subalpine riparian forests, four lower subalpine willow carrs, three upper subalpine willow carrs, and one subalpine wetland.     

Changes in riparian vegetation along the Colorado River and Rio Grande, Colorado

Changes in vegetation including area occupied, canopy cover, and maturity class of cottonwoods ( Populus spp.) within lower-elevation zones of the Colorado River and Rio Grande in Colorado were monitored over 25- and 27-year intervals, respectively, using photo-interpretative methods. Estimated loss of cottonwoods along the Colorado River was 2ha/km (-17.5%), and remaining stands had become more open and older. Cottonwoods along the Rio Grande increased 1.6 ha/km (9.3%) with minor canopy cover and maturity class changes. Area occupied by shrubs and river channel changed little along the Colorado River, but declined along the Rio Grande. Loss of hay meadow occurred along both rivers, whereas developed land increased along the Colorado River and farmland increased along the Rio Grande. Wildlife habitats along the Colorado deteriorated much more rapidly than those along the Rio Grande during monitored intervals.     

Elevational changes in woody vegetation along three streams in Washington County, Utah

Patterns of change in the woody vegetation (trees and shrubs) of the riparian communities of three streams draining the slopes of the Pine Valley Mountains in Washington County, Utah, were examined. Thirty-nine study sites were established adjacent to the streams along an elevational gradient where vegetation, plant population, and species data were taken. Vegetation and species patterns varied with respect to elevation and geographical location. Vegetation of the high- and low-elevation areas was distinctly different, while that of the mid-elevations was transitional to both. Geographical patterns showed a north-south transition in vegetation between the Mojave and Great Basin deserts. Size-class distribution curves for 11 tree species showed varying degrees of survival with respect to age and location. Root sprouting was highly important in the reproductive effort of six of the trees. Species distribution patterns followed transriparian and intrariparian gradients within the riparian community.     

Bird use of riparian vegetation along the Truckee River, Califonia and Nevada

The Truckee River in California and Nevada is subject to diverse water regimes and a corresponding variety of flow rates. Original riparian vegetation has been altered by these variable flow rates and by a variety of human uses resulting in loss of native riparian vegetation from its historic extent. We conducted bird surveys along the Truckee River during spring 193 to (1) determine relationships between birds and the present vegetation; (2) determine the importance of different vegetation types to sensitive bird species that have declined recently in the western United States due to competition from exotic plant species, cowbird ( Molothrus ater ) parasitism, reduction in nesting habitat, or other unidentified reasons; and (3) establish a monitoring program and collect baseline data for future comparisons. The most frequently detected bird species throughout the study was the Brown-headed Cowbird. The greatest number of bird species (98 of 116) was found in the native mixed willow ( Salix spp.) riparian scrub vegetation type. We recommend protecting the remaining native riparian vegetation types for bird habitat along the Truckee River.     

Classification of the riparian vegetation along a 6-km reach of the Animas River, southwestern Colorado

Riparian ecosystems are important components of landscapes, particularly because of their role in biodiversity. A first step in using a ""coarse-filter"" approach to riparian biodiversity conservation is to determine the kinds of riparian ecosystems. These ecosystems vary substantially in plant species composition along a single river reach, as well as between rivers, and yet the river-reach scale has received little attention. We sampled the vascular plant composition of 67 contiguous patches of riparian vegetation along the reach of the Animas River, in southwestern Colorado's San Juan Mountains, that is relatively undisturbed by human land uses. Using cluster analysis and detrended correspondence analysis, we identified eight riparian community types along the reach. Using a new technique, we combined overstory size-class data and understory cover data to identify community types. The eight community types, which are in part the products of past floods, are spatially arranged along the reach in relation to variation in valley morphology, tributary location, and geomorphic landforms. These eight community types do not necessarily represent successional stages of a single potential vegetation type. This study at the river-reach scale suggests that sampling and analysis, as well as conservation, may need to be turned to the scale of patchiness produced by flood disturbances in the riverine landscape, since vegetation varies significantly at this scale.     

Ecological impacts of Pogonomyrmex on woody vegetation of a Larrea-Ambrosia shrubland

Discovery of distinct mid-elevational bands of blackbrush ( Coleogyne ramosissima Torr.) shrublands on desert mountain slopes in the Mojave Desert caused an investigation of the relationships between environmental factors and Coleogyne distribution. Environmental factors were quantitatively examined to determine which were significant predictors of Coleogyne density at upper-elevational limits (ecotones) in the Spring Mountains of southern Nevada. Path analysis revealed significant, direct causal effects of air temperature, soil moisture, soil depth, and percent litter cover on the distribution of Coleogyne . Specifically, air temperature was a significant positive predictor, while soil moisture, soil depth, and percent litter cover were significant negative predictors of Coleogyne density, with the effects of other environmental variables parceled out. Path analysis also indicated that indirect effects of soil pH, bulk density, compaction, percent pore space, organic matter, soil temperature, salinity, cryptogam, and percent bare soil and rock cover on Coleogyne density were substantially more potent than their direct casual effects. Environmental attributes associated with elevational changes correlate with and may determine the density of Coleogyne shrubs at upper ecotones in southern Nevada.     

Establishment, growth, and early survival of woody riparian apecies at a Colorado gravel pit

Presence of a wetted edge during the period of seedfall was an effective predictor of suitable establishment (defined as germination and survival to the 1st autumn) locations for Populus deltoides subsp. monilifera, Salix amygdaloides, S. exigua, and Tamarix ramosissima seedlings during 3 successive years of a gravel pit revegetation project in Fort Collins, Colorado. At locations predicted to be suitable for establishment, position within the pit (possibly reflecting additional moisture provided by seepage) was a significant factor in determining whether establishment actually occurred. Cover of herbaceous species, which became established at the same time as, or after, woody seedlings, was positively related to probability of establishment for 8 of 11 species-year combinations, probably reflecting more favorable moisture conditions at certain locations. Herbaceous cover also was positively related to seedling height at the end of the 1st summer of growth for 9 of 11 species-year combinations. Neither establishment nor 1st-summer growth was consistently related to overall decline in the water table as estimated by the drop in surface-water level during the growing season. Flooding in the 1st spring after establishment was negatively related to subsequent survival for 5 of 8 species-year combinations. The 4 species established at different elevations in the pit, depending on location of the wetted edge during the period of seedfall, and there was no evidence that differential mortality subsequently altered their distribution along the elevation gradient. However, the primary objective in this study was to restore native woody species, and we attempted to maintain conditions conducive to meeting this objective. Differential postestablishment mortality may be more important in structuring the riparian community in more rigorous riverine environments.     

Small mammals within riparian habitats of a regulated and unregulated arid land river

In northwestern Colorado, flow regulation on the Green River has created a transitional plant community that features encroachment by upland vegetation into cottonwood ( Populus fremontii )-dominated, riparian forest on topographically high floodplain sites and reduced cottonwood regeneration on low floodplain sites. To assess how these changes might have affected small mammal distributions, in 1994 and 1995 we live-trapped during periods surrounding spring flooding at 3 sites: above and below the confluence of the regulated Green River and at the ecologically similar, but unregulated, Yampa River (reference site). More species were captured at the most regulated site along the Green River above its confluence with the Yampa River. Within sites, more species were captured in riparian habitats than adjacent upland habitats. Despite river regulation-induced habitat changes, we did not detect changes in species distributions within low and high floodplain habitat for Peromyscus maniculatus or Microtus montanus , but changes may have occurred for Dipodomys ordii . The total effect of regulation-induced habitat change on small mammal populations may not be fully revealed until current, mature cottonwood forests disappear and associated woody debris decomposes.     

Vegetation, soils, and hydrogeomorphology of riparian patch types of a dryland river

From a landscape perspective, riparian corridors can be viewed as mosaics of vegetation patches. We delineated 10 patch types within the floodplain of the San Pedro River (Arizona) on the basis of physiognomy, dominant overstory species, and tree size class; and we assessed differences in hydrogeomorphology, vegetation structure, plant species richness, and soil chemistry and texture. Patches of tamarisk ( Tamarix ), an introduced species, fell within the continuum of variation shown by other patch types in the landscape mosaic. Among the tree-dominated types, cottonwood-willow ( Populus-Salix ) and tamarisk patches were inundated more frequently than mesquite ( Prosopis ) patches, while cottonwood-willow patches had shallower groundwater than tamarisk or mesquite patches. Due to the wetter conditions, cottonwood-willow patches had a high relative abundance of wetland and exotic species in the understory. Tamarisk patches and wet shrublands ( Baccharis salicifolia—Salix exigua ) had high woody stem densities while cottonwood-willow patches had dense canopy cover. In association with differences in canopy cover, cottonwood-willow patches had low herbaceous species richness but high woody species richness, while tamarisk patches had high herbaceous and low woody species richness. Soil electrical conductivity, silt content, organic matter content, and available phosphorus increased from young to old stands of both tamarisk and cottonwood-willow, often resulting in greater differences between patches of different size/age class than between patches with different dominant species. Surface soil salinity (electrical conductivity) was low in all patches, including those dominated by tamarisk. Nitrate was abundant in soils of tamarisk patches (perhaps reflecting their high clay content) and wet shrubland patches. Dry shrublands ( Hymenoclea-Ericameria ) and wet shrublands were similar to young forest patches in having coarse soils with little organic matter.     

Monitoring temporal change in riparian vegetation of Great Basin National Park

Disturbance in riparian areas of semiarid ecosystems involves complex interactions of pulsed hydrologic flows, herbivory, fire, climatic effects, and anthropogenic influences. We resampled riparian vegetation within ten 10-m × 100-m plots that were initially sampled in 1992 in 4 watersheds of the Snake Range, east central Nevada. Our finding of significantly lower coverage of grasses, forbs, and shrubs within plots in 2001 compared with 1992 was not consistent with the management decision to remove livestock grazing from the watersheds in 1999. Change over time in cover of life-forms or bare ground was not predicted by scat counts within plots in 2001. Cover results were also not well explained by variability between the 2 sampling periods in either density of native herbivores or annual precipitation. In contrast, Engelmann spruce ( Picea engelmannii ) exhibited reduced abundance at all but the highest-elevation plot in which it occurred in 1992, and the magnitude of change in abundance was strongly predicted by plot elevation. Abundance of white fir ( Abies concolor ) individuals increased while aspen ( Populus tremuloides ) individuals decreased at 4 of 5 sites where they were sympatric, and changes in abundance in the 2 species were negatively correlated across those sites. Utility of monitoring data to detect change over time and contribute to adaptive management will vary with sample size, observer bias, use of repeatable or published methods, and precision of measurements, among other factors.     

Factors controlling structural and floristic variation of riparian zones in a mountainous landscape of the western United States

We examined landscape patterns in the physical conditions and vegetative composition of montane riparian zones to identify their most important sources of variation. Information on plant species cover and on physical characteristics that occur at coarse, medium, and fine scales was collected for 144 riparian plots located throughout the Lake Tahoe Basin, which straddles the California-Nevada border in the western United States. Constrained and unconstrained ordination analyses were used to identify the most important correlates of physical form and plant species composition. Through multivariate analysis of environmental variables (principal components analysis), vegetation data (detrended correspondence analysis), and the combined relationship between the environmental and vegetation data (canonical correspondence analysis), we consistently found that the greatest variation occurred along a gradient of decreasing valley width, decreasing stream sinuosity, and increasing stream slope. Although surface characteristics reflected a 2nd important source of variation in physical conditions, plant species distribution was not strongly correlated with riparian surface conditions. Strong correlations among physical variables that occur at different scales, such as between valley form and geofluvial surface and between geofluvial surface and surface conditions, support the use of a physically based hierarchical framework for organizing riparian zones within the landscape. Such a hierarchical framework would be useful for interpreting patterns in riparian structure and process at different scales and could be applied to riparian zones in other mountain landscapes of the western United States and elsewhere. Moreover, our finding that riparian plant species composition is most strongly correlated with environmental variables that occur at coarse to moderate scales, most of which can be derived from existing data, supports the idea that modeling montane riparian community distribution using topographic and remotely sensed data could be useful; however, a large degree of species variation, unexplained by the variables we collected, indicates that other variables, perhaps disturbance regime, should be included in such a venture. Nomenclature: Hickman (1993).     

Transverse pattern of vegetation on avalanche paths in the northern Rocky Mountains, Montana

Normal 0 false false false EN-US X-NONE X-NONE MicrosoftInternetExplorer4 /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Table Normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-priority:99; mso-style-qformat:yes; mso-style-parent:""; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:11.0pt; font-family:"Calibri","sans-serif"; mso-ascii-font-family:Calibri; mso-ascii-theme-font:minor-latin; mso-fareast-font-family:"Times New Roman"; mso-fareast-theme-font:minor-fareast; mso-hansi-font-family:Calibri; mso-hansi-theme-font:minor-latin; mso-bidi-font-family:"Times New Roman"; mso-bidi-theme-font:minor-bidi;} The pattern of vegetation on avalanche paths has usually been ascribed to the damage done by snowslides. In the northern Rocky Mountains the pattern of herbs, shrubs, and small trees appears to be more complex than could be accounted for by avalanche magnitude and frequency. The vegetation on one path in Montana illustrates that the topography of the path is a factor in the distribution of species. Three zones exist across avalanche paths: an inner zone of herbs and suffrutescent shrubs occupying a ravine, which is snow covered longer than elsewhere; flanking zones of dense shrubs and trees with flexible stems; and an outer zone of less dense shrubs that is more xeric. The pattern of vegetation seems to be due to avalanche-related stress rather than damage.        

Caddisfly (Trichoptera) assemblages along major river drainages in Arizona

Seventy-two caddisfly species in 36 genera and 15 families were collected along the Little Colorado, Verde, Gila, and Colorado rivers in Arizona. Brachycentrids, hydropsychids, limnephilids, and uenoids made up nearly 50% of the assemblage in forested sites above 2200 m. Hydropsychids, hydroptilids, and leptocerids made up 75% of the assemblage in grassland watersheds between 2200 and 1000 m, and 80% at sites below 1000 m in desert watersheds. Species richness averaged 16 at sites above 2200 m elevation, 7 between 2200 and 1000 m, and 3 below 1000 m. H' diversity was typically >3 at sites above 2200 m and <2.5 below 2200 m. Each river had distinct faunas likely due to isolation of drainages across semiarid landscapes. Low S&oslash;rensen similarity values were measured for caddisfly assemblages in the highly regulated Colorado River and the other 3 rivers. Only 17 species were collected at 14 sites along 700 km of the Colorado River. Indicator caddisfly species are provided for altered and less disturbed drainages. We added 7 species not previously reported in Arizona for a total of 154 caddisfly species in the state. We provide a baseline of caddisfly indicators for monitoring changes in ecosystem health during the predicted long-term drought and population expansion in southwestern USA.     

Genetic variation of woodrats ( Neotoma cinerea ) and deer mice ( Peromyscus maniculatus ) on montane habitat islands in the Great Basin

Seventeen loci were examined for polymorphism in four populations of Neotoma cinerea and Peromyscus maniculatus on isolated mountain ranges in the Great Basin, one population of each in the Sierra Nevada, and one of each in the Rocky Mountains. All Peromyscus populations had higher levels of heterozygosity than syntopic Neotoma populations.     

Hip glands in a natural population of montane voles ( Microtus montanus )

Average linear movement by populations of Dipodomys ordii, Microtus montanus, Perognathus parvus, and Peromyscus maniculatus &nbsp; was investigated over a 15-month period by live trapping on a low-level, radioactive waste disposal area in Idaho. No significant differences in movement among habitats were observed seasonally, excepting M. montanus in spring. Average linear movements within habitats ranged from 20 to 70 m for all species, but some patterns varied seasonally and among age classes for individual species. Although predation on contaminated small mammals from the disposal area is a vector of radionuclide transport, local movements by these rodents do not appear to be of sufficient magnitude to contribute significantly to redistribution of radioactive particles.     

Riparian vegetation along the middle Snake River, Idaho: zonation, geographical trends, and historical changes

A baseline study was conducted on an 83-km free-flowing reach of the Snake River between Swan Falls Dam and the Idaho-Oregon border. The research had 2 components: (1) field characterization and inventory of existing riparian flora, vegetation, and environment (soils, topography, streamflow), and (2) determination and mapping, using a geographic information system, of historic changes in riparian vegetation based on a time series (1938-39, 1957, 1969, 1987) of aerial photographs. The flora was diverse, with 185 species of vascular plants identified, 63 of which were exotics. Vegetation was structured vertically along the riverbank gradient into lifeform-defined habitat types: emergent, riparian shrub-forb, tree, transitional grass-shrub, and upland. Riverbank seepage, probably of agricultural origin, blurred zonation patterns on some sites and added species to the overall flora. Upstream-downstream differences existed in the physical characteristics and vegetation of river subreaches. Coverage of riparian woodland, island riparian and total riparian vegetation, and area of islands increased since the 1930s, with the greatest changes in the 1969-1987 interval. Possible contributing facts were (1) significant declines in annual minimum flows since the 1950s, (2) decreases in peak flows following the completion of Swan Falls Dam and some upstream dams since the 1920s, (3) introduction and spread of exotic tree species ( Elaeagnus angustifolia and Tamarix spp.) and (4) possible effects of intensive agriculture on river sediment load and soil nutrients. The introduction and proliferation of purple loosestrife ( Lythrum salicaria ) could have considerable future effects on vegetation-channel dynamics in the middle Snake River.     

Tree growth and regeneration response to climate and stream fow in a species-rich southwestern riparian forest

We studied the influence of climate variables, stream flow, and topography on regeneration and growth of several riparian tree species ( Acer negundo, Alnus oblongifolia, Fraxinus velutina, Juglans major, Platanus wrightii, Populus fremontii, Salix spp.) at an unregulated perennial stream, West Clear Creek, in central Arizona. A pulse of seedling regeneration occurred for Alnus, Fraxinus, Platanus, Populus , and Salix in 1995 and 1996 following high winter and spring surface flows in 1993 and high spring surface flow in 1995. In contrast, little regeneration occurred for Acer and Juglans under these conditions. Most seedlings occurred at the active channel topographic location, and few seedlings occurred at abandoned channel, gravel-boulder bar, and bench locations. Relationships between environmental variables and annual radial stem growth varied among species and between constrained and unconstrained reaches. High spring or winter surface flows were negatively related to growth of Acer, Alnus, and Platanus , whereas high spring surface flow was positively related to growth of Fraxinus . Positive relationships between precipitation and growth occurred only for Fraxinus and Juglans , suggesting greater use of surface soil water by these species. Annual radial growth was high for Platanus and Alnus, medium for Acer and Fraxinus, and low for Juglans . Overall, the tree species in our study responded individually, rather than collectively, in regeneration and growth to changes in stream flow and climatic variables.     

Effects of livestock grazing exlosure on aquatic macroinvertebrates in a montane stream, New Mexico

Aquatic macroinvertebrate populations inhabiting reaches of a stream within areas excluded from livestock grazing for a decade were markedly different from those in grazed areas when density, biomass, biotic condition indices, and mean chi square indices of the two populations were compared. Increased densities and biomasses of more tolerant forms of macroinvertebrates were observed in grazed reaches. Because pretreatment data were not available, differences in macroinvertebrate populations and relative tolerances of taxa in grazed and ungrazed areas could be as easily attributed to linear changes in stream habitat as to removal of domestic livestock. Results of this study have implications for the design of future research on the effects of livestock grazing on stream environments and biota: (1) baseline/pretreatment information is prerequisite, and (2) the study should take a watershed (ecosystem) approach.     

Winter site fidelity and body condition of three riparian songbird species following a fire

The effects of fire on nonbreeding songbird species in riparian habitat have not been studied. We compared body condition, within-year site fidelity, and between-year site fidelity of 3 songbird species ( Passerella iliaca , Fox Sparrow; Catharus guttatus , Hermit Thrush; and Regulus calendula , Ruby-crowned Kinglet) at 2 coastal riparian sites. Wildfire, which is rare in this habitat, had occurred at 1 of the sites before data collection. A significantly larger proportion of Passerella iliaca was recaptured in subsequent winters at the unburned site than at the burned site, but little difference was found between sites for Catharus guttatus or Regulus calendula . Body mass of all 3 species declined during winter at the burned site, but differences between sites were not significant. Similarly, body mass indices of new captures were lower at the burned site than the unburned site for all 3 species, but these differences were not significant. The within-year recapture rate for all 3 species combined declined at the burned site over the course of the study, possibly due to changes in vegetation structure caused by the fire. Overall, our data suggest that wintering songbirds were resilient to this disturbance, but that response to the post-fire environment differed among foraging guilds. Well-replicated studies that include pre-burn data are needed to evaluate the effects of this disturbance in riparian systems.     

Long-term vegetation dynamics in a cut western juniper woodland

Western juniper ( Juniperus occidentalis spp. occidentalis Hook.) expansion in the northern Great Basin has reduced shrubsteppe productivity and diversity. Chainsaw cutting of western juniper woodlands is commonly applied to remove tree interference and restore sagebrush plant communities. Studies assessing understory response following cutting have been limited to early successional stages and have not evaluated the effects of western juniper debris on plant succession. Cutting western juniper produces a large amount of debris which is commonly left on site, occupying a significant portion of treated areas. This study evaluated successional dynamics spanning 13 years after western juniper cutting. Four 0.45-ha blocks were selected on Steens Mountain in southeastern Oregon. Western juniper cover averaged 26% and mature tree density averaged 250 trees ? ha –1 . Blocks were cut in late summer 1991. Understory standing crop, cover, and density were compared among 3 locations: old canopy litter mats (canopy), interspace, and area underneath cut western juniper (debris). In the interspace, perennial grasses increased in cover and in standing crop relative to other functional groups. In canopy and debris locations, species composition shifted in the 6th year after cutting as annual grass cover, density, and standing crop increased. However, by 2003, perennial grass biomass was 2 times greater than annual grass biomass in canopy and debris locations. Because annual grasses increased in areas of debris accumulation, managers need to be cognizant of western juniper treatments that create safe sites that are favorable to the establishment of weedy species. Retaining western juniper debris on this site did not increase establishment and growth of perennial grasses compared to the interspace. A shift in perennial grass dominance from Thurber's needlegrass ( Achnatherum thurberianum [Piper] Barkworth) to bottlebrush squirreltail ( Elymus hystrix [Nutt.] Smith) occurred in areas of debris accumulation. Our results demonstrated that long-term vegetation evaluations are necessary to properly assess management activities and disturbance.