Karyotypes of four Artemisia species: A. carruthii, A. filifolia, A. Frigida, and A. spinescens

Artemisia carruthii and A. frigida of the subgenus Artemisia and A. filifolia and A. spinescens of the subgenus Dracunculus all have chromosome numbers based on x = 9. Diploid (2n = 18) karyotypes of each species are composed of large, medium, and small chromosomes that are mainly metacentric and submetacentric. The individual karyotypes are similar but distinctive. Artemisia filifolia ’s karyotype and chemistry is quite similar to that of Section Tridentatae, but A. filifolia has significant morphological differences with respect to the Tridentatae. Artemisia spinescens includes a tetraploid (2n = 36) population as well as diploid populations. Karyotypic analysis of a tetraploid A. spinescens suggests that it is an autotetraploid, thus carrying out a common theme in Artemisia (autopolyploidy).     

The parasitic wasp genera Skiapus,Hellwigia, Nonnus,Chriodes, and Klutiana (Hymenoptera,Ichneumonidae): Recognition of the Nesomesochorinae stat. rev. and Nonninae stat. nov. and transfer of Skiapus and Hellwigia to the Ophioninae

The ichneumonid wasp genera Skiapus Morley, Hellwigia Gravenhorst, Nonnus Cresson, Chriodes Förster, and Klutiana Betrem are each formally removed from the Campopleginae on the basis of morphological, molecular, and simultaneous analyses using PAUP* and the optimization alignment program POY. Skiapus and Hellwigia are shown to form a monophyletic group with the Ophioninae to which they are transferred. Nonnus is treated as comprising the Nonninae stat. nov. and Chriodes and Klutiana (sometimes treated as a junior synonym of Chriodes) are treated as comprising the Nesomesochorinae stat. rev. The status of Nesomesochorinae and Nonninae is not fully resolved as they are not consistently recovered as separate groups, forming a single clade in some analyses. We keep them separate because of this uncertainty. Molecular synapomorphies within the D2–3 expansion region of the 28S rDNA gene show the utility of this gene region in determining subfamily‐level placement within the higher Ophioniformes.