Geography of exotic plants adjacent to campgrounds, Yellowstone National Park, USA

Eleven campgrounds in Yellowstone National Park were studied to determine the geography of 10 specific exotic plant species adjacent to campgrounds. Exotics were found in only 6 campgrounds. Six species were found at Mammoth campground, a low-elevation, dry site with year-round use. Only 2 species were found in the other 5 campgrounds. Exotics decreased with distance from Mammoth campground out to 6 m and then increased, suggesting a spread in their distribution. Significant associations were found between exotic presence and both open and closed canopies and low levels of disturbance. Generally, exotics decreased with an increase in cover of other vegetation forms. Five species were found most frequently in big sagebrush habitat types.     

Arbuscular mycorrhizae in thermal-influenced soils in Yellowstone National Park

Mycorrhizae are common plant-fungal symbioses occurring in most land plants. Despite their ubiquity, little is known about the distribution of arbuscular mycorrhizae (AM) in extreme environments. We surveyed for the presence of AM in thermal sites in Yellowstone National Park (YNP) where soils are characterized by extreme pHs, elevated temperatures, and toxic element concentrations. Plants at 5 sites, growing in soils with rooting-zone temperatures up to 48°C and soil pH values as low as 3.4, were mycorrhizal (colonization levels from 4% to 34%). Soils from a sparsely vegetated thermal area and an adjacent, continuously vegetated transition area differed significantly in rooting-zone temperature (35°C vs. 26°C), acidity (pH 3.8 vs. 5.4), electrical conductivity (2.22 vs. 0.49 mmhos cm -1 ), Fe (181.3 vs. 48.5 mg kg -1 ),Mn (7.2 vs. 98.2 mg kg -1 ), and Zn (2.3 vs. 4.5 mg kg -1 ). Mycorrhizal infectivity potential (MIP) was 77% greater in the transition soils, with colonization levels of 26% and 46% in thermal and transition soils, respectively. Furthermore, colonization of Agrostis scabra , Dichanthelium lanuginosum , and Mimulus guttatus was found to be consistently high throughout the growing season (from 48% to 72%). It is possible that AM are essential for plant life on the edge of thermal areas, and that either or both symbionts are specifically adapted to their environment. Further research is required to elucidate AM function in and specific adaptations to YNP's thermal areas.     

Annotated checklist of exotic vascular plants in Yellowstone National Park

Documentation of the arrival of exotic vascular plants in Yellowstone National Park has been sporadic. An annotated checklist of exotic vascular plants is presented, with information about the approximate arrival time in the park of each species and the current extent of the infestation. Yellowstones flora includes 187 exotic vascular plant species (14.8% of the flora), and the park has an extrapolated mean number of exotic species per 10 km 2 of 47.3. The situation in Yellowstone is compared with other areas in North America. The increase in exotics mirrors a corresponding increase in visitation.     

Managing a complex exotic vegetation program in Yellowstone National Park

The number of documented exotic plants in Yellowstone National Park has increased from 85 known in 1986 to over 185 today. Exotic plants are substantially impacting the parks natural and cultural resources and are a high management priority. We have adopted an integrated weed management approach with regard to exotic vegetation, emphasizing prevention, education, early detection and eradication, control, and, to a lesser degree, monitoring. The program involves over 140 staff with program expenditures averaging approximately $190,000 annually. Prevention actions include requiring approved gravel on construction projects; banning hay in the backcountry and allowing transport of only certified weed-seed-free hay through Yellowstone; requiring construction equipment to be pressurecleaned prior to entering the park; and native species revegetation after road, housing, and other construction projects have disturbed ground. Over 4500 acres, primarily along roadsides and in developed areas, are surveyed annually in early detection efforts with emphasis placed on eradicating small, new infestations of highly invasive species such as sulfur cinquefoil ( Potentilla recta L.) and leafy spurge ( Euphorbia esula L.). Control efforts focus on about 30 priority species, such as spotted knapweed ( Centaurea maculosa Lam.), oxeye daisy ( Chrysanthemum leucanthemum L.), and hoary cress ( Cardaria draba [L.] Desv.) using chemical, mechanical, and cultural techniques. A total of 2027 acres were treated during 1998, whereas control efforts for 12 species occurred on 2596 acres during the previous 3-year period, 1995-1997. Strong and expanding partnerships with other federal, state, and local agencies and private companies contribute to management efforts within the park. Future program goals emphasize increases in base funding to ensure continued weed management efforts as well as expanding survey, monitoring, and reclamation efforts. Ultimately, a more rigorous assessment of program effectiveness is desired.     

Bottom-up factors influencing riparian willow recovery in Yellowstone National Park

After the elimination of wolves ( Canis lupis L.) in the 1920s, woody riparian plant communities on the northern range of Yellowstone National Park (YNP) declined an estimated 50%. After the reintroduction of wolves in 1995–1996, riparian willows ( Salix spp.) on YNP’s northern range showed significant growth for the first time since the 1920s. However, the pace of willow recovery has not been uniform. Some communities have exceeded 400 cm, while others are still at pre-1995 levels of <80 cm mean height. We took intensive, repeated measurements of abiotic factors, including soil and water-table characteristics, to determine whether these factors might be contributing to the varying pace of willow recovery. Willows at all of our study sites were “short” (<250 cm max. height) prior to 1995 and have recovered to varying degrees since. We contrasted “tall” (>250 cm max. height) willow sites where willows had escaped elk ( Cervus elaphus L.) browsing with “short” willow sites that could still be browsed. Unlike studies that manipulated willow height with fences and artificial dams, we examined sites that had natural growth differences in height since the reintroduction of wolves. Tall willow sites had greater water availability, more-rapid net soil nitrogen mineralization, greater snow depth, lower soil respiration rates, and cooler summer soil temperatures than nearby short willow sites. Most of these differences were measured both in herbaceous areas adjacent to the willow patches and in the willow patches themselves, suggesting that they were not effects of varying willow height recovery but were instead preexisting site differences that may have contributed to increased plant productivity. Our results agree with earlier studies in experimental plots which suggest that the varying pace of willow recovery has been influenced by abiotic limiting factors that interact with top-down reductions in willow browsing by elk.     

Effects of browsing by native ungulates on the shrubs in big sagebrush communities in Yellowstone National Park

The effects of elk ( Cervus elaphus ), pronghorn ( Antilocapra americana ), and mule deer ( Odocoileus hemionus ) browsing on shrubs in big sagebrush ( Artemisia tridentata ) communities were monitored over a 31-year period in Yellowstone National Park. Ungulates were restricting Wyoming big sagebrush (spp. wyomingensis ) heights, size, and recruitment on the lower-elevation stratum only, while no such suppression was observed on the high-elevation stratum. Parallel increases in mountain big sagebrush (spp. vaseyana ) densities and cover occurred over the study period on both browsed and unbrowsed sites at the higher-elevation stratum, although big sagebrush, green rabbitbrush ( Chrysothamnus viscidiflorus ), and horsebrush ( Tetradymia canescens ) were slightly taller and crown sizes were slightly larger on unbrowsed than browsed sites. Wyoming big sagebrush utilization (percent leader use) was eight times higher (￣ x = 87 ± 7.2% by pronghorns, mule deer, and elk) on the low-elevation winter ranges stratum (the Boundary Line Area [BLA] portion of the winter range), while mostly mountain big sagebrush with leader use averaged only 11 ± 4.1% (nearly all by elk) on the high-elevation range stratum. In addition, annual aboveground biomass production of big sagebrush did not differ between browsed and unbrowsed study sites on the high-elevation stratum of the winter range. Population turnover was higher on browsed plots versus unbrowsed plots. No difference was observed in percent dieback of big sagebrush adult plants between browsed and unbrowsed plots at the higher stratum. Browsing did not influence the number of leaves or seedstalks per plant ( P > .05), but leaves averaged 45% longer and seedstalks 42% longer on browsed big sagebrush. Ungulate browsing, however, apparently suppressed production, germination, and survival of Wyoming big sagebrush on the low-elevation stratum. Numbers of Wyoming big sagebrush declined 43% and cover declined 29%, 1957-1990, on browsed sites on the BLA. Annual biomass production on browsed sites at the low-elevation stratum was only 6-35% that of unbrowsed sites, and big sagebrush recruitment was less on browsed sites. Percent leader use of big sagebrush did not differ between the period of ungulate reductions, 1962-1969, and the 1980s on the lower stratum (￣ x = 87% leader use), but utilization was less on higher portions of the winter range during the period of elk reductions (￣ x = 2%) than during the 1980s following cessation of elk controls (￣ x = 11%).     

Chemical and biological characteristics of desert rock pools in intermittent streams of Capitol Reef National Park, Utah

Chemical variability and biological communities of rock pools found in small desert drainage basins of Capitol Reef National Park were characterized over 8 mon in 1994. Neither flooding, drying, nor the presence or absence of surrounding vegetated wetlands had a great effect on chemical composition, which was very dilute and fluctuated somewhat in response to rain events. Neither flooding nor drying affected the composition of biological communities in the pools. Summer storms affected only a few drainages at a time, and only a few study pools of significant volume dried completely during the hot, dry summer. This suggests that only a portion of the Waterpocket Fold aquatic community is ever disturbed at a time, leaving undisturbed areas as a source of recovery. Pools bordered by vegetated wetlands always supported greater numbers of species throughout the year than those bordered only by bedrock, but the same taxa were found in both vegetated and bedrock pools. The rock pool fauna in Capitol Reef National Park appear to be resilient to climatic variability.     

Macroinvertebrate richness is lower in high-elevation lakes vs nearby streams: evidence from Grand Teton National Park,Wyoming

Mountain ecosystems will be strongly impacted by climate change, yet little is known of extant biodiversity in high-elevation lakes, particularly in North America. In this study, we sampled the littoral zone of six alpine and subalpine lakes in Grand Teton National Park (GRTE), Wyoming, to characterise invertebrate diversity and environmental variation in these climate change-threatened ecosystems. Overall, we collected 19 aquatic invertebrate taxa, and found that each lake harboured a unique assemblage of invertebrates despite close geographic proximity in some instances (e.g. less than 5 km). The results of this study complement previous efforts focused on macroinvertebrate diversity of streams in the Teton Range, highlighting much lower diversity in montane lakes vs nearby streams. Taken together, the two studies establish an important baseline understanding of mountain freshwater biodiversity in GRTE. With rapidly changing hydrologic inputs to mountain lakes driven primarily by the recession of alpine glaciers, these results may help target aquatic invertebrates to monitor as climate change affects the region. Moreover, these data clarify habitat factors, both biotic and abiotic, that influence high-elevation lake assemblages of the Teton Range.     

Macroinvertebrate community structure and related environmental variables in two forks of the Virgin River, Utah

Using multivariate analysis techniques, we analyzed variation in the structure of the benthic macroinvertebrate community in the North Fork and East Fork of the Virgin River in and above Zion National Park in concert with a number of naturally occurring and anthropogenic environmental variables. Correspondence analysis revealed that the community structure in the 2 streams was markedly different despite occurring in areas similar in topography, elevation, and terrestrial vegetation cover. Subsequent analyses of the 2 streams separately using canonical correspondence analysis to test for strength of association of environmental variables with variations in benthic macroinvertebrate community structure revealed that communities in the East Fork of the Virgin River were more associated with nitrogen-related variables influenced by agricultural and urban activities along the river and that communities in the North Fork were most strongly associated with naturally occurring physical variables.     

Effects of 2,4-D on a Populus tremuloides community in the western United States-22 years after treatment

Normal 0 false false false EN-US X-NONE X-NONE MicrosoftInternetExplorer4 st1\:*{behavior:url(#ieooui) } /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Table Normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-priority:99; mso-style-qformat:yes; mso-style-parent:""; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:11.0pt; font-family:"Calibri","sans-serif"; mso-ascii-font-family:Calibri; mso-ascii-theme-font:minor-latin; mso-fareast-font-family:"Times New Roman"; mso-fareast-theme-font:minor-fareast; mso-hansi-font-family:Calibri; mso-hansi-theme-font:minor-latin; mso-bidi-font-family:"Times New Roman"; mso-bidi-theme-font:minor-bidi;} Quaking aspen ( Populus tremuloides Michx.) stands were accidentally sprayed with 2,4-D in a sagebrush control program in western Wyoming in 1958. We visited the site during the summer of 1981 to evaluate the long-term effect on the aspen trees and the associated vegetation. Initially, some observers believed that the aspen had been “destroyed.” Subsequent data indicate just the opposite—aspen stocking appeared to have been improved by the treatment. On two of the sampled clones, 22 years after spraying, there were approximately 17,000 more suckers/ha on the sprayed than on the unsprayed plots. These are adequate numbers to restock the site to pretreatment densities. Although undergrowth vegetation appeared to be changed as a result of the treatment, this cannot be attributed solely to the herbicide because heavy grazing, mostly by domestic livestock, has occurred on the grazing allotment. A similarity index was calculated between sprayed and unsprayed portions of the same aspen clones. Forbs were still less on the sprayed areas, whereas grasses were similar on the sprayed and unsprayed areas. Spraying apparently does not have as adverse an effect on aspen communities as some conservationists originally thought.        

Distribution of the subterranean amphipod Stygobromus in central Colorado streams, with notes on the interstitial community

As part of a larger study on the aquatic life of streams in central Colorado, efforts were made to determine the distribution of interstitial organisms, with emphasis on the subterranean amphipod Stygobromus . In a preliminary screening-level study, sampling for Stygobromus was conducted in 1985 at 47 sites along the Front Range of Colorado at elevations generally matching the type locale for S. coloradensis and S. pennaki at the confluence of the North Fork of the South Platte River and its mainstem. A more intensive follow-up study was conducted in 1988 at 30 sites concentrating on the upper South Platte River basin. In the follow-up study both hyporheic and shore zone (phreatic) habitats were sampled using a variety of techniques (Bou-Rouch pump, Karaman-Chappuis technique, and a coarsemesh wide-mouth net, as appropriate). In the preliminary study Stygobromus spp. were collected at only 12 of 47 study sites, with all collections in the vicinity of the type locale. In the follow-up study Stygobromus were found at 16 of 30 sites sampled, extending the known distribution in Colorado of S. coloradensis and S. pennaki well beyond the type locale. The use of multiple sampling techniques was important in locating these organisms. In addition to the amphipod Stygobromus , sampling sites in the follow-up study contained a rich interstitial community, including copepods, bathynellids, tardigrades, archiannelids, and ostracods.     

Habitat and plant distributions in hanging gardens of the Narrows, Zion National Park, Utah

Hanging gardens are insular plant communities of the Colorado Plateau. This study examines hanging gardens in the Narrows, Zion National Park, Utah. The floristic similarity of gardens and the presence of species in classes of variables which characterize the habitat are disclosed. Although the gardens are isolated and dissimilar, the individual species are not restricted in the range of habitat found. Floristic dissimilarity cannot be attributed to differences in habitat. These results are compared to studies of hanging gardens in eastern Utah.       

Flood frequency and the assemblage of dispersal types in hanging gardens of the Narrows, Zion National Park, Utah

Hanging gardens of the Narrows, Zion National Park, Utah, are plant communities growing at permanent seeps on the canyon walls. The gardens are isolated from each other and from other plant communities by vertical expanses of sandstone. Gardens consist mostly of herbaceous species less than 1 m tall. Though not individually species-rich, the hanging gardens are diverse as a group, and very dissimilar. This study considers two explanations of the heterogeneous distribution of species in hanging gardens. The assemblages of dispersal types in gardens of different spatial attributes and disturbance frequencies are examined. The G H statistic, a log likelihood ratio test, analyzes the incidence of dispersal types among classes of three spatial and two disturbance variables. The disturbance variables of expected flood frequency and soil depth segregate dispersal types; and the spatial variables of area, distance to possible seed sources, and relative isolation do not. Ferns and mosses, dispersing through spores, dominate a heterogeneous fugitive guild in the flood-prone gardens. Infrequently flooded gardens support more large-seeded species.     

At the crossroads from Asia to Europe: spring migration of raptors and black storks in Dadia National Park (Greece)

Migrating raptors and black storks (Ciconia nigra) were studied in Dadia National Park, Greece, during spring 2003–2005. Vantage points and transects were used to evaluate magnitude, phenology, local variation and direction. We observed 23 species and 2030 individuals, including 715 common buzzards (Buteo buteo), 547 black storks and 136 short-toed eagles (Circaetus gallicus). Species-specific migration peaks were detected, starting in the second half of March, e.g. common buzzard, short-toed eagle, black stork, and ending early May, e.g. Levant sparrowhawk (Accipiter brevipes), red-footed falcon (Falco vespertinus), honey buzzard (Pernis apivorus). Most raptors were observed in the Evros valley, which may function as a migration corridor. The average direction of passing raptors was north. Species’ proportions and phenology in the study area were similar to those in neighbouring Bosphorus and Marmara Flyways. Further migration monitoring should be established in the area, which will provide important information not least to inform wind farm development location.     

Habitat use by the Persian onager,Equus hemionus onager (Perissodactyla: Equidae) in Qatrouyeh National Park,Fars, Iran

Iran’s Persian onager populations are critically endangered. This study of their natural history in Qatrouyeh National Park provides insights for enhancing their conservation. The population as a whole is greatly affected by weather. Wind, rain and cold drive populations from the plains to the valleys of hill-valley habitats. Vegetation features and water also influence habitat use, but differently for different sex and reproductive classes. Females with juveniles use plains with high-quality vegetation, whereas females without young and solitary territorial males choose those of intermediate quality. Females with young foals are also found closest to watering points. Future translocation of Persian onagers will only succeed if prospective habitats have sufficient hill-valley refuges and enough plains with winds to moderately hot conditions. Sufficient plains supporting high-quality vegetation near water for lactating females must co-exist with plains of moderate-quality vegetation that attract females without young, so reducing crowding and competition.     

Short-term effects of wildfire on montane stream ecosystems in the southern Rocky Mountains: one and two years post-burn

The frequency and intensity of wildfire in the western United States has increased over the last century, creating a heterogeneous mosaic of landscapes in various stages of recovery. The 2002 Hayman Fire was one of the largest wildfires in Colorado history and was unprecedented for its speed and intensity, with over 50%–70% of the burn area classified as moderate to high severity where much of the canopy crown was consumed. We evaluated the short-term impact of the Hayman Fire on ecological properties in montane stream ecosystems in the summers of 2003 and 2004, one and two years post-fire. Fire significantly altered surface temperature, dissolved oxygen concentration, and the chemical composition of stream water, including concentrations of nitrate, phosphate, and mineral salts. However, fire did not have significant impacts on stream conductivity, pH, or total concentrations of cations or anions during our study period. Streams in the burn area contained fewer benthic macroinvertebrate taxa compared to unburned streams during the year after fire and contained lower invertebrate densities and biomass compared to reference streams 2 years post-fire. Average C:N ratio of the benthic macroinvertebrate community was significantly and negatively related to stream nitrate concentration, possibly due to a shift in community composition or invertebrate nitrogen acquisition in fire-affected streams.     

Deer mouse, Peromyscus maniculatus, and associated rodent fleas (Siphonaptera) in the arctic-alpine life zone of Rocky Mountain National Park, Colorado

Normal 0 false false false EN-US X-NONE X-NONE MicrosoftInternetExplorer4 /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Table Normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-priority:99; mso-style-qformat:yes; mso-style-parent:""; mso-padding-alt:0in 5.4pt 0in 5.4pt; mso-para-margin:0in; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:11.0pt; font-family:"Calibri","sans-serif"; mso-ascii-font-family:Calibri; mso-ascii-theme-font:minor-latin; mso-fareast-font-family:"Times New Roman"; mso-fareast-theme-font:minor-fareast; mso-hansi-font-family:Calibri; mso-hansi-theme-font:minor-latin; mso-bidi-font-family:"Times New Roman"; mso-bidi-theme-font:minor-bidi;} Peromyscus maniculatus and related small rodents have been examined for ectoparasites in the tundra region of the Rocky Mountain National Park 1974–1979. One hundred and ninety-four P. maniculatus were examined from two tundra sites. Flea infestation rates were 1.9 fleas per mouse examined and 4.1 fleas per infested mouse. Species taken in significant numbers were Monopsyllus thambus (51 percent), Peromyscopsylla hesperomys (34 percent), Malaraeus euphorbi (9 percent), and Catallagia calisheri (4 percent). Peromyscus maniculatus host/flea relationships in the tundra are compared with those in other life zones in the park.       

A comparison of the structure and function of the cerotegument in two species of Cryptostigmata (Acarina)

Cerotegument plastron structure and function are compared in two species of non-related Cryptostigmata from different families in two environments prone to flooding, namely soil and leaf litter, as represented by Euzetes globulus (Nicolet) (Euzetidae), and the intertidal zone by Ameronothrus marinus (Banks) (Ameronothridae). Differences in cerotegument structure reflect the degree of exposure to mechanical stresses and hydrostatic pressure encountered in these two environments. Cerotegument plastrons communicate with the tracheal system in heavily armoured species, e.g. E. globulus, but gas transfer may occur across the cuticle in those species having a lightly sclerotized leathery cuticle, e.g. A. marinus.     

Life cycle of the net-spinning caddisfly, Cheumatopsyche analis (Banks) (Trichoptera: Hydropsychidae), in two small Front Range urban streams, Fort Collins, Colorado

The life cycle of Cheumatopsyche analis (Banks) is described from 2 urban streams located in north central Colorado. Larvae were collected every 15 days during spring, summer, and fall and every 30 days in winter from August 1999 through August 2000. A total of 8652 larvae were collected, and head capsules were measured to the nearest 0.01 mm. The life cycle of C. analis at both sites appears to be partially bivoltine with a long, almost continuous period of recruitment after peak emergence in June. Some offspring from the early generation that emerges in May probably emerge as adults in September, while the remainder of the population overwinter as instars III through V. Life cycle patterns for C. analis in these 2 urban streams were most similar to other populations of Cheumatopsyche from Virginia.