A survey of the genus Geometra Linnaeus (Lepidoptera,Geometridae, Geometrinae)

The type genus of the family Geometridae, Geometra Linnaeus, is reviewed. In addition to the 16 species known worldwide, two new species are described: Geometra neovalida sp. nov. from China, and Geometra burmensis sp. nov. from Burma (Myanmar). All the known species are redescribed. Genitalia of all species are described and illustrated, most for the first time. Generic characters based on all species are summarized. The monophyly of the genus and its relationship with allied genera are reviewed and discussed in detail. Most species fall into two species groups, named the papilionaria and smaragdus groups, on the basis of the morphology of the male genitalia. Biological information is provided where available. Illustrations of moths and genitalia are presented.     

Taxonomic review of a new genus,Diathera gen. n., from Southeast Asia (Lepidoptera,Geometridae: Larentiinae)

Diathera gen. n., a new larentiine genus from central and southwestern China, is described. Three new species are named and described: fluctuata sp. n., metacolorata sp. n. and brunneata sp. n. All species and their genitalia are described and illustrated. A key is provided and the monophyly of the genus and its relationships to the allied taxa are briefly discussed.     

A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae)

The genus Pandalus Leach, 1814, is revised based upon the abundant material from collections in various museums or institutions in the world. Altogether 20 species are recognized which appear to form five groups. Nineteen species are recognized in Pandalus, three of which are described as new: P. curvatus from southern Japan, and P. chani and P. formosanus from Taiwan. Four informal species groups are also recognized within the genus: P. montagui group (P. montagui Leach, 1814; P. borealis Krøyer, 1838; P. goniuruStimpson, 1860; P. jordani Rathbun, 1902; P. tridens Rathbun, 1902; and P. eous Makarov, 1935); P. stenolepis group (P. stenolepis Rathbun, 1902; P. curvatus sp. nov.); P. hypsinotus group (P. hypsinotus Brandt, 1851; P. danae Stimpson, 1857; P. prensor Stimpson, 1860; P. gracilis Stimpson, 1860; P. gurneyi Stimpson, 1871; P. nipponensis Yokoya, 1933; P. teraoi Kubo, 1937; P. chani sp. nov.; and P. formosanus sp. nov.); and P. platyceros group (P. platyceros Brandt, 1851; and P. latirostris Rathbun, 1902). The P. platyceros group appears to be most closely related to the genus Pandalopsis Bate, 1888, but Pandalus is retained as a possible paraphyletic group. Protandrous hermaphroditism is known in all but two species of the genus (P. curvatus and P. formosanus), for which only specimens of either male or female have been available. Pandalus propinqvus G. O. Sars, 1870, is transferred to a new monotypic genus Atlantopandalus, because of its lack of hermaphroditism and possession of some unique morphological characters, including one indicating a close relationsip to Dichelopandalus Caullery, 1896. All species are fully described and illustrated. The affinities and important morphological variations of the species are discussed. A key for adults is presented for the identification of the species. Biogeography of the genus is briefly discussed.     

A review of the genus Talavera Peckham and Peckham, 1909 (Araneae,Salticidae)

The species of the Holarctic jumping spider genus Talavera are reviewed. Five new species are described: Talavera ikedai sp. n. (¦ §) from Japan and Korea; T. krocha sp. n. (¦ §) from France, Ukraine and Kyrghyzstan; T. parvistyla sp. n. (¦ §) from central and northern Europe; T. sharlaa sp. n. (§) from Tuva; and T. tuvensis sp. n. (¦) from Tuva. A lectotype is designated for Euophrys monticola Kulczy ` ski, 1884. A key to the 14 species now encompassed in this genus, and distributional maps, are provided for all of them. Arguments are given for placing the names Euophrys poecilopus Thorell, 1873 and Attus westringi Simon, 1868 as nomina dubia. Talavera aperta Miller, 1971 is removed from synonymy with both Talavera monticola (Kulczy ` ski, 1884) and Talavera thorelli (Kulczy ` ski, 1891).     

A revision of the Asian and European species in the subgenus Amiota Loew (Diptera,Drosophilidae) and the establishment of species-groups based on phylogenetic analysis

A total of 53 Amiota (s. str.) species (75% of the world total) from Asia and Europe, including 10 new species, kamui, kimurai and planata from Japan, and aristata, cuii, macai, magniflava, nuerhachii, spinata and watabei from China, are reviewed with designation of two new synonymies and phylogenetic analysis. Based upon the result of cladistic analysis with 31 adult male morphological characters, the following conclusions are deduced: (1) The subgenus Amiota is monophyletic, so far as the Asian and European forms are concerned. (2) Seven monophyletic groups are recognized within this subgenus. Two of them correspond to known species-groups, the apodemata and sinuata groups, and the remaining five are established as new species-groups, the nagatai, basdeni, taurusata, alboguttata and rufescens groups. A key to all the studied species from Asia and Europe is provided.     

Oribatid mites of Liebstadia (Acari: Oribatida: Scheloribatidae) from Mongolia,with notes on taxonomy of the genus

Three new species of oribatid mites belonging to the genus Liebstadia are described from Mongolia. The new species are characterized by the poorly developed pteromorphae; the club-shaped head of the sensilli; the absence of the dorsosejugal suture; the dorso-ventral thickness, and the length and arrangement of notogastral setae. In addition, L. similis (Michael, 1888) is redescribed and recorded for the first time in Mongolia. Protoribates serratomarginatus Mahunka, 1983 is synonymized with L. similis. A key to the known species of the genus Liebstadia is given.     

A revision of Heterosignum (Asellota,Isopoda, Paramunnidae) with descriptions of three new species

Three new species of Heterosignum are described from Japan. Heterosignum elegans sp. nov. differs from its congeners in having stout preorbital lobes on the head, very long lateral spine-like processes and eyestalks, and very long article 2 of male pleopod 2. Heterosignum ohtsukai sp. nov. is distinctive in having a marked stout mid-dorsal spine on the head, long mid-dorsal spines on the pereonites, and article 2 of antenna 1 with a distolateral protrusion. Heterosignum otsuchiensis sp. nov. has single lateral spine-like processes on pereonites 1-7 as in H. ohtsukai, but is distinguished from it in having short mid-dorsal spines on pereonites 1-7 and long basis of pereopod 1. Heterosignum mutsuensis Gamô, 1976 is redescribed using newly collected specimens. Coulmannia unicornis Kensley, 1976 is assigned to Heterosignum after careful examination of the types. The diagnosis of Heterosignum is emended to include all the five species so far recorded.     

A revision of Caridina yunnanensis and its allied species (Crustacea: Decapoda: Caridea: Atyidae) from Yunnan,southern China,with description of one new species

The taxonomy of the Caridina yunnanensis and its allied species (Caridea: Atyidae) from Yunnan Province in southern China, is revised. A neotype for C. yunnanensis Yu, 1938, is designated. Additional information is provided on C. disparidentata Liang, Yan and Wang, 1984; C. dianchiensis Liang and Yan, 1985; C. mongziensis Liang, Yan and Wang, 1987; and C. disjuncta Cai and Liang, 1999. One new species, Caridina impensa, is described. A key to members of the species group is included.     

A review of the north-western Pacific species of the genus Paguristes (Decapoda: Anomura: Diogenidae), I. Five species initially reported by Ortmann (1892) from Japan

Five species of the hermit crab genus Paguristes Dana, 1851, initially reported by Ortmann (1892) from Japan, are revised based on the materials reported by Ortmann (1892), Doflein (1902), Balss (1913) and Miyake (1978), and on newly collected specimens. The specific status of P. palythophilus Ortmann, 1892, is clarified, and a species confused with P. palythophilus is described as new, P. albimaculatus. It has been revealed that P. kagoshimensis Ortmann, 1892, is actually a junior subjective synonym of P. digitalis Stimpson, 1858. The species heretofore assigned to P. kagoshimensis is described as new, P. versus. It has been confirmed that Paguristes acanthomerus Ortmann, 1892, is a valid species and that P. barbatus sensu Ortmann, 1892, is the same species as P. ortmanni Miyake, 1978. Further, a new species, P. doederleini, is described for P. setosus sensu Ortmann, 1892. It is also shown that a part of the group of specimens identified as Paguristes puniceus by Miyake (1978) is referable to P. doederleini. In addition to the three new species recognized herein, the four previously described species are also redescribed in detail in order to make clear their taxonomic status.     

A new genus of viviparous gyrodactylid (Monogenea) from the Great Barrier Reef,Australia with descriptions of seven new species

Acanthoplacatus gen. nov., a new genus of viviparous gyrodactylid, is described from the fins and skin of siganid fishes from the Great Barrier Reef, Australia. The genus is characterized by a muscular, tube-like haptor with 16 marginal hooks on the posterior margin. The ventral lobe of the haptor is located anteriorly relative to the dorsal lobe and contains a pair of hamuli and a ventral bar with posteriorly-projecting ventral bar membrane. A dorsal bar is absent. Five pairs of posterior gland cells surround the posterior terminations of the gut. The male copulatory organ is a muscular, non-eversible bulb with several spines around the distal opening. Species of Acanthoplacatus have a bilateral excretory system consisting of six pairs of flame cells and a pair of excretory bladders. Seven new species are described: Acanthoplacatus adlardi sp. nov. and A. amplihamus sp. nov. from Siganus punctatus (Forster, 1801), A. brauni sp. nov. from S. corallinus (Valenciennes, 1835), A. parvihamus sp. nov. from S. vulpinus (Schlegel and Mueller, 1845), A. puelli sp. nov. from S. puellus Schlegel, 1852, A. shieldsi sp. nov. from S. lineatus (Valenciennes, 1835) and A. sigani sp. nov. from S. fuscescens (Houttuyn, 1782). Species can be discriminated by shape and size of the hamuli, marginal hooks and ventral bar and by male copulatory organ sclerite morphology. Three species (A. brauni sp. nov., A. shieldsi sp. nov. and A. sigani sp. nov.) were assessed for seasonal variation of sclerite size. Ten of thirteen morphological characters showed seasonal variation in size for at least one of the species. The characters were longer in winter except dorsal root tissue cap width. Only one character, marginal hook length, showed significant seasonal variation for all three species. Species of Acanthoplacatus were observed to attach using only the marginal hooks and the role of hamuli in attachment is unclear. The dorsal fin of the host is the preferred site for most species but the anal fin, caudal fin and body surfaces are preferred by some species. Prevalences for species range from 57 to 100%.     

Two new species of protandric pheretimoid earthworms belonging to the genus Metaphire (Megascolecidae: Oligochaeta) from Taiwan

This paper describes two new species, each consisting of two subspecies, of protandric, octothecal pheretimoid earthworms from Taiwan. They are Metaphire paiwanna sp. nov. typica, Metaphire paiwanna var. liliumfordiI subsp. nov., Metaphire bununa sp. nov. typica and Metaphire bununa var. glareosus subsp. nov. The var. typica of both new species occurred on the west side of the Central Mountain Range, whereas the subspecies, liliumfordi and glareosus, occurred on the east side of the range. Apparently, the Central Mountain Range constitutes a geographical barrier for isolation and subspeciation. Metaphire paiwanna has a male disc medial to male pore (opening of copulatory pouch) with a horizontal ridge, the character shared by holandric and sexthecal Amynthas swanus (Tsai) of Taiwan and Metaphire aggera (Kobayashi) of Korea and Manchuria, whereas M. bununa resembles more closely holandric, octothecal Metaphire riukiuensis (Ohfuchi) of the Riukiu Islands than protandric, octothecal Metaphire stephensoni (Michaelsen) of Indochina.     

A revision of the Caridina serrata species group,with descriptions of five new species (Crustacea: Decapoda: Caridea: Atyidae)

The taxonomy of the Caridina serrata species group (Caridea: Atyidae) is revised. Neotypes of C. serrata Stimpson, 1860, and C. cantonensis Yu, 1938, are designated. Additional information on Caridina sphyrapoda Liang and Zhou, 1993 is provided. Five new species of the Caridina serrata species group are described, namely Caridina nanaoensis from Guangdong Province, C. apodosis from Hong Kong, and C. yulinica, C. wumingensis and C. mutata from Guangxi Province, all in southern China. A key to all members of the species group is included.     

A study of the Exoprosopa busiris (Jaennicke) group of Bezzi (1924) in tropical Africa (Diptera: Bombyliidae)

The tropical African species of the genus Exoprosopa Macquart placed by Bezzi (1924) in his group 10, E. busiris are reviewed. A key, short diagnoses and outline distributions are provided for each species. E. albonigra Bezzi and E. saskae Szilády are shown to be synonyms of E. luteicosta Bezzi; E. engyoptera Hesse a synonym of E. brevinasis Bezzi; and E. ferrieri Hesse a synonym of E. decastroi Hesse. Two new species are described; E. selenops from Kenya and E. glossops from Nigeria.     

A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins

The eumedonid genera Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847, are revised. Members of both genera are obligate symbionts with sea urchins. Eumedonus is separated from Gonatonotus mainly by the presence or absence of crests on the merus of the ambulatory legs. Eumedonus, as here defined, contains five species, viz. E. niger H. Milne Edwards, 1834 (type species), E. vicinus Rathbun, 1918, E. zebra Alcock, 1895, E. brevirhynchus n. sp., and E. intermedius n. sp. Gonatonotus, as here re-diagnosed, includes three species, viz. G. pentagonus White, 1847 (type species), G. granulosus (MacGilchrist, 1905), n. comb. and G. nasutus n. sp.     

A revision of the genus Ceratomerus (Diptera: Empididae: Ceratomerinae) of New Zealand

The genus Ceratomerus Philippi (Diptera: Empididae; Ceratomerinae) of New Zealand is revised and a key to species given. Thirteen species are recognized and the following new species are proposed: C. biseriatus sp. nov., C. flavus sp. nov., C. earlyi sp. nov., C. melaneus sp. nov., C. brevifurcatus sp. nov., C. tarsalis sp. nov. and C. vittatus sp. nov. The biology and phylogeny are briefly discussed.     

Notes on Bombyliidae (Diptera) from the southern borderlands of the Sahara with descriptions of new species

The paper is a review of the genus Pellenes (s. lat.) in the fauna of Central Asia and the Caucasus. The genus Pellenes(s. lat.) is redefined and subdivided into four subgenera according to the structure of genitalia: Pellenes(s. str.), Pelpaucus subgen. nov., Pelmultus subgen. nov. and Pelmirus subgen. nov. Altogether 23 species have been treated, of which nine are described as new to science: Pellenes amazonka sp. nov. (female), P. badkhyzicus sp. nov. (male), P. borisi sp. nov. (female), P. karakumensis sp. nov. (male), P. bonus sp. nov. (male), P. pamiricus sp. nov. (male and female), P. pseudobrevis sp. nov. (male and female), P. sytchevskayae sp. nov. (male and female) and P. turkmenicus sp. nov. (male and female). Five species are newly synonymized: P. maderianus Kulczyński, 1905 with P. epularis (O. P.-Cambridge, 1872); P. kulabicus Andreeva, 1976 and P. simoni (O. P.-Cambridge, 1872) with P. geniculatus (Simon, 1868); and P. albomaculatus Peng and Xie, 1993 with P. denisi Schenkel, 1963; P. tauricus (Thorell, 1875) is removed from synonymy with P. simoni (O. P.-Cambridge, 1872) and is shown to be a junior synonym of P. nigrociliatus (Simon in L. Koch, 1875). A lectotype is designated for Pellenes brevis Simon, 1868. P. campylophorous (Thorell, 1875) is recognized to be nomen dubium. Pellenes calvus (Simon, 1868) is excluded from the Central Asian fauna of Pellenes. Evarcha lapponica (Sundevall, 1832), comb. nov. (ex Pellenes), is proposed. Some morphological terms, e.g. the median septum, the epigynal flaps, the central blindending pocket and the newly proposed ‘compound terminal apophysis’, are discussed in relation to their homology.     

A taxonomic revision of the Oriental spittlebug genus Paphnutius Distant (Homoptera: Cercopidae)

The spittlebug genus Paphnutius is transferred from the Tomaspidini to the Considiini. The genus is revised and redescribed to contain nine species. A key to species is given and all species are redescribed and illustrated with their distributional data provided. One new species is described and one new generic synonymy, six new specific synonymies and four new combinations are established. Five lectotypes are newly designated. Paphnutius costimaculus Metcalf and Horton is considered as a species incertae sedis.     

Two new phoxocephalids (Crustacea: Amphipoda: Phoxocephalidae) from the south-west Atlantic

A new genus and species, Parafoxiphalus longicarpus, and a new species, Metharpinia protuberantis belonging to the family Phoxocephalidae, are described from the Argentine Sea. Both taxa were collected from sandy substrata in the intertidal and subtidal regions. Parafoxiphalus longicarpus is related to Foxiphalus Barnard, 1979 and they share many characters, most of them in plesiomorphic condition. Metharpinia protuberantis is an intermediate species between Metharpinia Schellenberg, 1931 and Microphoxus Barnard, 1960, and its assignation to the first genus is discussed.     

A revision of the new riodinid butterfly genus Pseudotinea (Lepidoptera: Riodinidae)

A revision of the new South American riodinid genus Pseudotinea is presented that includes taxonomic and ecological notes, distributional data and illustrations of adults and genital structures for all species. The systematic position of Pseudotinea within the Riodinidae, its ecology and biogeography are discussed. Five species are recognized: volcanicus Callaghan and Salazar, hemis Schaus and caprina Hewitson are transferred from Calydna (comb. n.) and two new species, P. eiselei and P. gagarini, are described here. The genus is confined to the Andes and south-east Brazil where all species are very rare and many endangered.