Contribution to the systematic revision of the African cyprinid fish genus Labeo: species from the Lower Zaire river system

The species of the genus Labeo from the Lower Zaire river system have been revised on the basis of 284 preserved fishes, including type material of 18 nominal species cited or described from this region. Morphometrics, meristic counts, morphological observations and radiographs have been used for this study. All results obtained are supported by statistical analyses. From the 18 nominal species reported, only 12 species are recognized as valid. Two major groups were identified: one group has rows of conical papillae on the inner surface of lips; and the other group presents rows of transverse plicae on the inner surface of lips. The first group has two species: L. lineatus and L. weeksii. One synonymy is proposed: L. lineatus (partim) = L. weeksii. In the second group, 10 species have been identified: L. longipinnis, L. degeni, L. barbatus, L. macrostoma, L. falcipinnis, L. cyclorhynchus, L. parvus, L. greenii, L. nasus and L. sorex. One unidentified taxon apparently represents a species new to science, but this must be confirmed by a wider revision of the genus which is the subject of further research. Some synonymies in this group are proposed: L. brichardi (partim) = L. barbatus; L. lineatus (partim) = L. degeni. Other synonymies already established by Reid (1985), have been confirmed: L. lividus = L. barbatus; L. brichardi (partim) = L. macrostoma; L. variegatus = L. cyclorhynchus. One synonymy has been restored: L. velifer = L. longipinnis. Finally, one synonymy remains in question: L. chariensis = L. parvus. A key to species is provided.     

First record of the hyperparasite Liriopsis pygmaea (Cryptoniscidae,Isopoda) from a rhizocephalan parasite of the false king crab Paralomis granulosa from the Beagle Channel (Argentina), with a redescription

The Cryptoniscidae are epicaridean isopod parasites or hyperparasites of other crustaceans. Liriopsis Schultze in Müller, , one of the genera included in this family, now contains two nominal species: L. pygmaea (Rathke, ) and L. monophthalma (Fraisse, ). Both of these species infest rhizocephalan cirripeds, which are in turn parasites of hermit crabs. Among the false king crabs, Paralomis granulosa (Jacquinot, 1847), captured commercially in the Beagle Channel in 1996–1998, we found 31 specimens of the rhizocephalan Briarosaccus callosus Boschma, 1930 infested by one or more specimens of L. pygmaea. Neither L. pygmaea nor L. monophthalma has been reported previously for the southern seas. Although unidentified isopod hyperparasites have been found on B. callosus infesting other lithodids from Crozet Islands, South Georgia Island and Canadian Atlantic waters, this is the first time that one of these hyperparasites has been identified as a member of the genus Liriopsis. Since the differences between L. pygmaea and L. monophthalma remain obscure, the epicaridium and cryptoniscus larvae and three females stages of L. pygmaea are herein described from the material collected in the Beagle Channel.     

Hitchhiking with the Vikings? The anthropogenic bumblebee fauna of Iceland – past and present

The number of bumblebee species found in Iceland has doubled from three to six in 35 years. This paper considers the colonisation of Iceland by bumblebees; updates information on Bombus(Pyrobombus) jonellus (Kirby), B. (Megabombus) hortorum (L.) and B. (Bombus) lucorum (L.); presents findings for two new colonists, B. (Pyrobombus) hypnorum (L.) and B. (Thoracobombus) pascuorum (Scopoli); and considers the arrival of another species, B.(Pyrobombus) pratorum (L.), that may have become established. The context of bumblebee dispersal to remote islands is considered. Many of the restricted number of insect species that have colonised North Atlantic islands have done so in association with man. B. (Bombus) terrestris (L.) has been used as a pollinator in glasshouses in Iceland since 1994, and it may also have become naturalised. Barcoding is needed to check the identity of some specimens. B. jonellus has probably been present for at least several hundred years; genetic studies should help determine whether the timescale for its arrival is compatible with an anthropogenic origin. It is suggested that consideration is given to ways of reducing both unintended introduction of bumblebee species with cargo, and the release and disease risk of those imported intentionally for commercial use.     

Phorbas posidoni n.sp. (Porifera: Poecilosclerida) from the Aegean Sea,with a discussion of the family Anchinoidae

A new species of the genus Phorbas Duchassaing and Michelotti, 1864 (junior synonym Anchinoe Gray, 1867) is described from the Aegean Sea (Eastern Mediterranean). Phorbas posidoni n.sp. is distinguished from the known Mediterranean species of Phorbas primarily by its elaborate repent-ramose habit combined with an extensive, spongin-enforced, plumoreticulate skeleton of oxea-like tornotes and the usual echinating acanthostyles and arcuate chelae. As in most Phorbas the surface bears characteristic areolae. The only other elaborate Mediterranean species, P. mercator (Schmidt) differs clearly in spiculation, since it lacks microscleres and has strongyles as the main skeletal elements. Phorbas fictitius (Bowerbank), P. tenacior (Topsent) and P. paupertas sensu Topsent are all incrusting and differ in spicule form and sizes as well. Likewise, Phorbas species from the neighbouring East Atlantic each show clear differences with P. posidoni n.sp. The genus Phorbas is discussed and compared with Pronax Gray (1867) sensu Lévi (1973). It is proposed to unite the two genera because the alleged sharp difference between the two (only oxea-like tornotes in the skeletal tracts in Phorbas, only acanthostyles in the skeletal tracts in Pronax) is compromised by a range of intermediate conditions in various species.     

Neotropical Lispe (Diptera: Muscidae): notes,redescriptions and key to species

The following species of the fly genus Lispe are redescribed: L. albitarsis (Stein, 1898), L. latana Snyder, 1949, L. levis (Stein, 1911), L. lisarba Snyder, 1949, L. nasoni (Stein, 1898), L. serotina Wulp, 1896, L. setuligera (Stein, 1911) and L. vilis (Stein, 1911). Additionally, a new diagnosis is proposed for L. rufitibialis Macquart, 1843, and in order to fix the identity of the name the lectotype is designated. To facilitate species identification, a pictorial key for the species of Lispe in the Neotropical region and distribution maps are presented, together with the first published illustrations of the male terminalia of L. latana, L. levis and L. lisarba, and the female of L. lisarba.     

Recent and fossil Lunulitidae (Bryozoa,Cheilostomata) 6. Lunulites sensu lato and the genus Lunularia from Australasia

The characters of the genus Lunulites Lamarck, and of some closely related genera are described and discussed. The genus Lunularia Busk is reintroduced, and three species from Australasia are described. One species, L. parvicella, is known only as an Australian Tertiary fossil, but the remaining two, L. capulus and L. repandus, have an extensive Tertiary-to-Recent Australasian distribution. L. capulus is apparently confined to Australian waters and deposits, but L. repandus has now been found in the Miocene and Recent of New Zealand, as well as off the southern and western Australian coasts.     

Two new genera of Bryozoa (Calloporidae) from New Zealand

Two new genera of the bryozoan family Calloporidae are described from New Zealand. The first, Leptinatella, is introduced for L. gordoni n. sp., specimens of which have been referred in the past to Watersia militaris (Waters) but are distinct from this species, which is a phase of the cribrimorph Corbulipora tubulifera (Hincks). The second genus, Bryocalyx, is introduced for B. cinnameus n. sp., which has small, conical colonies anchored by rhizoids. Another species of Bryocalyx is known from a few fragments only: and has also been referred in the past to Watersia sp. It is briefly discussed, but left unnamed here.     

Taxonomic revision of the genus Littorophiloscia Hatch, 1947 (Crustacea,Isopoda, Oniscidea) with description of six new species

In the present revision 15 species of Littorophiloscia are recognized. Six species, namely L. tropicalis, L. koreana, L. amphindica, L. pallida, L. bifasciata and L. hawaiiensis are described as new. Littorophiloscia isabellae (Vandel, 1973) is considered to be a junior synonym of L. albicincta (Vandel, 1973), and Philoscia dartevellei Brian, 1953 and Vandeloscia orientalis Roman, 1977 are considered to be junior synonyms of L. culebrae (Moore, 1901). Alloniscus compar sens. Vandel, 1952 nec Budde-Lund, 1893 is considered to be distinct from L. riedli (Strouhal, 1966) and it has been renamed L. tropicalis sp. nov. The genera Bilawrencia Vandel, 1973, Helenoscia Vandel, 1977, Vandeloscia roman, 1977 and Sayoscia Schultz, 1983 are considered to be synonymous with Littorophiloscia. Diagnostic characters of all the species are illustrated. A map and comments on the distribution of the 15 species are included.     

New and little‐known species of Didemnidae (Ascidiacea,Tunicata) from Australia (Part 3)

The majority of the 58 species discussed, including 10 new species, were collected by scuba divers at 5–10?m in waters around Australia. Species are from Leptoclinides (11), Polysyncraton (11), Didemnum (19), Trididemnum (7), Lissoclinum (8), Clitella (1), and Diplosoma (1), and new species are in all except Trididemnum and Diplosoma. Additional characters (including a pyloric vesicle reported previously in the Holozoidae and several unrelated didemnid taxa) have been detected for the monotypic genus Clitella Kott, , which is recorded for only the second time. A review of known Australian ascidian species confirms the Didemnidae as the most speciose ascidian family in these waters. In this family, there appears not to be appreciable gene flow between tropical and temperate waters and few species have a continuous tropical–temperate range. A preponderance of Western Pacific non‐indigenous species is in the north, while indigenous species, some probably isolated from related tropical ones, are dominant in the temperate waters of the southern half of the continent. Although intraspecific variation and convergence obscures species differences, some aspects of the living organisms detected in in situ photographs contribute to identification. Keys to Australian didemnid species described since the publication of the Australian Ascidiacea part 4, Didemnidae (Kott ) are included.     

New bariliin cyprinid fishes from West Africa,with a consideration of their biogeography

Five new species of bariliin cyprinid fishes are described from West Africa. Three belong to the genus Leptocypris and two to Raiamas. One species of each genus is from Guinea (Konkoure river) and from Sierra Leone (Taia and Little Scarcies rivers), the other, a Leptocypris, is from Cameroon (Cross River). A taxon of indeterminate generic identity from Guinea is also described. The phylogenetic relationships of the bariliins are hypothesized and in this light their geographical distribution is compared with that of other cladistically analysed African freshwater fish groups. The significance to West African fish distribution of the Togo-Benin (Dahomey) Gap is discussed.     

New species of Eretmocerus Haldeman (Hymenoptera: Aphelinidae) parasitizing Crenidorsum turpiniae (Takahashi) and Aleurothrixus floccosus (Maskell) (Hemiptera: Aleyrodidae) from Taiwan

A key to females of 11 Eretmocerus species occurring in Taiwan is provided. Two new species, E. tongxiaoensis Shih & Polaszek, sp. nov. and E. lannae Shih & Polaszek, sp. nov. found infesting Crenidorsum turpiniae (Takahashi) and Aleurothrixus floccosus (Maskell) respectively, are described. Five more Eretmocerus species are recorded here as new to Taiwan: E. bisetae Hayat, E. flavus Krishnan & David, E. queenslandensis Naumann & Schmidt, E. rui Zolnerowich & Rose and E. trialeurodis Hayat. New host records for four of these Eretmocerus species from five whitefly species are presented. The whitefly Aleurothrixus floccosus (Maskell) is recorded for the first time from Taiwan.     

The Andean tarantulas Euathlus Ausserer, 1875, Paraphysa Simon, 1892 and Phrixotrichus Simon, 1889 (Araneae: Theraphosidae): phylogenetic analysis,genera redefinition and new species descriptions

Based on a phylogenetic analysis we revised the Chilean tarantula genera Euathlus and Paraphysa. As results of our analyses, within a wider context of South American lineages, Paraphysa is synonymous of Euathlus, and Phrixotrichus is resurrected. Euathlus and Phrixotrichus are sister genera, supported by their shared palpal organ morphology, especially at the ventral position of the distal prolateral inferior keel, also by spermathecal receptacles with a lateral chamber, and tarsal claws without teeth. Both genera are redefined based on cladistic results, and some of their constituent species are transferred. We describe four new species, and for the first time present the spermathecae of Euathlus parvulus comb. nov. All species described for both genera are diagnosed and keyed. Euathlus now includes: Euathlus antai Perafán and Pérez-Miles sp. nov., Euathlus atacama Perafán and Pérez-Miles sp. nov., Euathlus condorito Perafán and Pérez-Miles sp. nov., Euathlus manicata (Simon 1892) comb. nov., Euathlus parvulus (Pocock, 1903) comb. nov. and Euathlus truculentus L. Koch, 1875. Phrixotrichus now comprises: Phrixotrichus jara Perafán and Pérez-Miles sp. nov., Phrixotrichus scrofa (Molina, 1788) comb. nov. and Phrixotrichus vulpinus (Karsch, 1880) comb. nov. Furthermore, Paraphysa riparia Schmidt and Bolle, 2008 is synonymized with Eupalaestrus weijenberghi (Thorell, 1894), Paraphysa pulcherrimaklaasi Schmidt, 1991 is transferred to Maraca Pérez-Miles, 2006 and Paraphysa peruviana Schmidt, 2007 is considered a nomen dubium.

http://zoobank.org/urn:lsid:zoobank.org:pub:62B49343-DCF0-4AFE-8154-19F9D50E9AA0     

Anomuran Crustacea (Paguridea,Porcellanidae, and Hippidea) from Oman,principally from Dhofar province,southern Oman

Anomura (Crustacea, Decapoda) are described from 5 collection sites in Dhofar (southern Oman) and from near Muscat (northern Oman). These consist of the following: Paguridea: Aniculus erythraeus, Calcinus (3 species), Cestopagurus coutieri, Clibanarius (3 species), Dardanus (5 species), Diogenes (2 species), Paguristes (2 species) and Coenobita scaevola; Porcellanidae: Pachycheles (2 species), Petrolisthes (7 species), Pisidia (3 species) and Polyonyx (1 species, probably undescribed); and Hippidea: Emerita holthuisi, Hippa pacifica and Albunea steinitzi. The anomuran fauna of the region is not well known, and that of Dhofar has not previously been investigated. The present records establish a number of significant range extensions. The anomura species composition of Dhofar is discussed in relation to that of other areas of the Indian Ocean. In general, pagurideans appear less cosmopolitan, and more parochial within the region than porcellanids. This is discussed in relation to larval dispersal and other factors.     

Three new species of Cecidophyinae (Acari: Eriophyidae) from China

Three new eriophyoid mite species (Acari: Eriophyoidea: Eriophyidae) from China are described and illustrated. They are Cecidophyes truncatis sp. nov. on Acer truncatum Bunge (Aceraceae), Cecidophyes hirsutes sp. nov. on Cardamine hirsute L. (Cruciferae) and Paracolomerus fopingacer sp. nov. on Acer sp. L. (Aceraceae). All the new species described herein are vagrants on the host plant.     

Gastrointestinal helminths from fifteen species of Litoria and Nyctimystes (Anura: Hylidae) from Papua New Guinea

Three hundred and ninety‐nine individuals representing 10 species of Litoria and five species of Nyctimystes (Hylidae), L. arfakiana, L. bibonius, L. eucnemis, L. genimaculata, L. impura, L. infrafrenata, L. modica, L. nasuta, L. thesaurensis, L. wollastoni, N. gularis, N. kubori, N. papua, N. pulcher and N. semipalmatus collected in Papua New Guinea were examined for helminths. Gravid individuals representing one species of Monogenea (Parapolystoma bulliense), one species of Digenea (Mesocoelium monas), 18 species of Nematoda (Aplectana krausi, Aplectana macintoshi, Aplectana zweifeli, Cosmocerca novaeguineae, Falcaustra papuensis, Icosiella papuensis, Maxvachonia adamsoni, Maxvachonia ewersi, Meteterakis crombiei, Moaciria moraveci, Ochoterenella papuensis, Oswaldocruzia bakeri, Parapharyngdon maplestoni, Parathelandros allisoni, Parathelandros andersoni, Physalopteroides milnensis, Rhabdias australiensis, Seuratascaris numidica) and one species of Acanthocephala (Acanthocephalus bufonis) were found. Also included were immature forms representing one species of Cestoda (as a metacestode), three species of Nematoda (Abbreviata sp., Rictulariidae gen. sp., Ascarididae gen. sp. in cysts), one species of Acanthocephala (unidentified cystacanth), and one species of Pentastomida [Raillietiella sp. (nymph)]. The mean helminth species richness for infected hosts was 1.3±0.6 SD (range one to three species). Sixty‐seven new host records and two new locality records are reported.     

Three new endogean species of Laparocerus Schönherr, 1834 from the Canary Islands (Coleoptera,Curculionidae)

Three new species of the weevil genus Laparocerus Schönherr (1834) adapted to the edaphic environment of the Canary Islands are described: L. edaphicus n. sp., L. oromii n. sp. and L. lopezi n. sp. They live in the humid laurel forests of Tenerife (Anaga Mountains), La Gomera (Garajonay National Park) and Gran Canaria (Valsendero), respectively. The small size and somewhat tubular body shape is a common character among them, as is the reduction of the eyes, which are totally absent in L. oromii n. sp. Despite these common adaptive traits, the three species seem not to be directly related. A key for the identification of all the known cryptozoic (edaphic and troglobitic) species of Laparocerus is provided.     

The cytogenetics of Lotus (Leguminosae)

Chromosome numbers for 108 species of the genus Lotus (Leguminosae) are now known. This represents an increase of 32 new species which have been reported since a chromosome atlas for Lotus was published in 1965. At this rate, with just over one chromosome number being reported for a new species per year, it will require almost a century for the chromosome numbers to be determined for all of the species in the genus. A review of the cytogenetic features which have been observed in the genus through karyotype analyses, chromosome banding, Feulgen cytophotometry, anther culture, HCN reaction and interspecific hybridization is presented. These include cytomixis and B chromosomes (both found in hybrids of diploid species in the L. corniculatus group when L. alpinus was one of the parents), desynapsis, binucleate cell formation, polytene chromosomes in suspensor cells and trisomics in L. pedunculatus. There has been a reduction of HCN content in the species with evolutionary development which is correlated with basic chromosome number and geographic distribution. Classical karyotype analyses may not be a suitable method to investigate the parentage of L. corniculatus because of the chromosomal repatterning which has occurred during the evolutionary development of the closely related diploid species.     

Helminths of ten species of Litoria frogs (Anura: Hylidae) from Papua New Guinea

One hundred and ninety-one frogs collected in Papua New Guinea from 10 species of the genus Litoria, namely, L. angiana, L. arfakiana, L. caerulea, L. congenita, L. eurynastes, L. gasconi, L. infrafrenata, L. iris, L. kumae and L. modica, were examined for helminths. Eighteen species of helminths were found: one species of Monogenea, Parapolystoma bulliense; three species of Digenea, Diplodiscus amphichrus, Halipegus zweifeli and Mesocoelium monas; 13 species of Nematoda, gravid specimens of Aplectana macintoshii, Cosmocerca novaeguineae, Falcaustra papuensis, Maxvachonia adamsoni, Maxvachonia flindersi, Maxvachonia ingens, Meteterakis crombiei, Moaciria moraveci, Parathelandros mastigurus, Physalopteroides milnensis, Pseudorictularia dipsarilis, Seuratascaris numidica, and larvae (in cysts) of Abbreviata sp. and, one species of Acanthocephala, Pseudoacanthocephalus bufonis. Mean number of helminth species per host species was 3.0 ± 1.4 SD (range 1–5), mean number of helminth species per infected frog was 1.1 ± 0.4 SD (range 1–2), and mean number of helminths per infected host was 33.3 ± 5.8 SD (range 1–36). Twenty-eight new host records and two new locality records are reported. The biogeography of the recovered helminths is discussed.     

Zoeal stages of Hiplyra variegata (Rüppell, 1830) (Crustacea: Brachyura: Leucosiidae) reared in the laboratory and collected from plankton at Al-Kharrar creek,central Red Sea

The zoeal stages of Hiplyra variegata are described and illustrated for the first time. Zoea I was obtained from an ovigerous female, and the zoea II and zoea III stages were captured in the plankton (their identity was confirmed by rearing some of them from earlier stages). Comparisons are made with the only other species of the genus of which larvae are known, i.e. H. platycheir. The presence of a lateral spine on the carapace of zoea I is one of the most important characters that can be used for easy recognition of H. variegata. It is important to note that the third zoea of H. variegata has four small spines at each posterolateral margin of the carapace, in contrast to three spines in other species.