A Chengjiang-type fossil assemblage from the Hongjingshao Formation （Cambrian Stage 3） at Chenggong,Kunming, Yunnan

A new Chengjiang-type fossil assemblage is reported herein from the lower part of the Hongjingshao Formation at Xiazhuang village of Chenggong,Kunming,Yunnan.The fossil assemblage,named as Xiazhuang fossil assemblage,yields predominantly soft-bodied fossils,including arthropods,brachiopods,priapulids,lobopods and some problematic taxa,with arthropods being the most dominant group.Preservation and composition of the fossil assemblage are very similar to the typical Chengjiang biota,which is preserved in the middle Yu’anshan Formation in the large area of eastern Yunnan.The associated trilobites demonstrate that the soft-bodied fossil assemblage belongs to the late Qiongzhusian in age(Stage 3,Cambrian),suggesting that the Hongjingshao Formation is probably a diachronous lithostratigraphic unit ranging from the upper Qiongzhusian to the lower Canglangpuan stages in eastern Yunnan.The fossil assemblage from the Xiazhuang area fills up the missing link between the typical older Chengjiang biota and the younger Malong and Guanshan biotas,making eastern Yunnan a unique area in the world to reveal the early evolutionary history of animals and palaeocommunity dynamics during the‘‘Cambrian explosion’’.     

Modern optics in exceptionally preserved eyes of Early Cambrian arthropods from Australia

Despite the status of the eye as an "organ of extreme perfection", theory suggests that complex eyes can evolve very rapidly. The fossil record has, until now, been inadequate in providing insight into the early evolution of eyes during the initial radiation of many animal groups known as the Cambrian explosion. This is surprising because Cambrian Burgess-Shale-type deposits are replete with exquisitely preserved animals, especially arthropods, that possess eyes. However, with the exception of biomineralized trilobite eyes, virtually nothing is known about the details of their optical design. Here we report exceptionally preserved fossil eyes from the Early Cambrian (～ 515 million years ago) Emu Bay Shale of South Australia, revealing that some of the earliest arthropods possessed highly advanced compound eyes, each with over 3,000 large ommatidial lenses and a specialized 'bright zone'. These are the oldest non-biomineralized eyes known in such detail, with preservation quality exceeding that found in the Burgess Shale and Chengjiang deposits. Non-biomineralized eyes of similar complexity are otherwise unknown until about 85 million years later. The arrangement and size of the lenses indicate that these eyes belonged to an active predator that was capable of seeing in low light. The eyes are more complex than those known from contemporaneous trilobites and are as advanced as those of many living forms. They provide further evidence that the Cambrian explosion involved rapid innovation in fine-scale anatomy as well as gross morphology, and are consistent with the concept that the development of advanced vision helped to drive this great evolutionary event.     

淮南地区新元古代刘老碑组叠层石消失原因初探

安徽淮南地区新元古代刘老碑组主要由页岩和中薄层泥质、粉砂质灰岩组成。在该组上部中薄层砂质灰岩中产有该地区新元古代最古老的叠层石Baicalia和?Linella。大量的臼齿碳酸盐脉体穿插在该组叠层石的基本层中的现象较为独特,该组上部发育的滑塌构造和臼齿碳酸盐岩脉体的特征表明该沉积时期发生过强烈的地震活动;此外,在叠层石礁体周围还有大量的风暴沉积产出。这些现象表明,该组叠层石发育于风暴活动和地震活动交织的环境之中,地震活动是导致该组叠层石消失的主要原因。     

Comparative study of Cambrian lobopods <Emphasis Type="Italic">Miraluolishania</Emphasis> and <Emphasis Type="Italic">Luolishania</Emphasis>

The rare fossil Miraluolishania described by Liu et ah from the Lower Cambrian Chengjiang Lagerstatte in 2004 is regarded as an arthropod sphinx because it bears mosaic features of both Iobopods and arthropods. The discovery of this rare transitional form offers direct fossil evidence for exploring the relationship between Iobopods and arthropods. However, some scientists consider Miraluolishania to be a junior synonym of Luolishania because the former superficially resembles the latter in general appearance. Considering the significant differences between the two taxa, a thorough comparative study of Miraluolishania and Luolishania leads to the conclusion that there are definitely two different genera. Nevertheless, the ＂Luolishania＂ of the Haikou area is indeed ＂Miraluolishania＂, whereas Luolishania is most likely the typical genus of the Maotianshan area of Chengjiang County.     

Primitive deuterostomes from the Chengjiang Lagerst?tte (Lower Cambrian, China).

Cambrian fossil-Lagerst?tten (sites of exceptional fossil preservation), such as those from Chengjiang (Lower Cambrian) and the Burgess Shale (Middle Cambrian), provide our best window into the Cambrian 'explosion'. Such faunas are known from about 40 localities, and have yielded a widely disparate series of taxa ranging from ctenophores to agnathan fish. Recent excavations of the Chengjiang fossil-Lagerst?tte, known from a series of sites near Kunming in Yunnan, south China, have resulted in the discovery of several new forms. In conjunction with material described earlier, these provide evidence for a new group of metazoans, the vetulicolians. Several features, notably a series of gill slits, suggest that this group can throw light on an early stage of deuterostome diversification.     

澄江动物群国家地质公园科学价值及旅游研究

澄江动物群国家地质公园是一个以保护具有全球意义的、实证"寒武纪生命大爆发"及地球早期生命起源和演化的澄江动物群及其赋存载体地层的非常珍稀的以古生物化石群为主题的国家级地质公园.主要以科考科普性旅游为主,具有不可替代、无与伦比的科学价值.针对澄江动物群国家地质公园的旅游特点,对旅游开发中存在的优劣势进行分析,对如何公园如何开展旅游进行研究.     

The earliest-known ancestors of Recent Priapulomorpha from the Early Cambrian Chengjiang Lagersttte

It was suggested and eventually has been profferedthat the representatives of all extant phyla, including someminor group, occurred during the Cambrian radia-tions[1—10]. Many priapulid-like fossil worms have beenreported from Cambrian Lagerst?tten, such as the EarlyCambrian Chengjiang Lagerst?tte[11—16], the Middle Cam-brian Burgess Shale[17], and the Middle Cambrian Kailifauna[18]. The palaeoscolecidan worms from these Lager-st?tten have been regarded as either a stem group of thepri…     

A venomous arthropod in the Early Cambrian Sea

A number of recently evolved animals possess poison glands for feeding and/or defense.However,examples of such animals are rare in the fossil record.We report a fossil arthropod Isoxys curvirostratus from the Early Cambrian Chengjiang biota of China.This species is regarded as the oldest known venomous arthropod based on the presence of venomous glands in its head region.The adult animal is 2-5 cm long and the body is covered entirely with a carapace.The presence of large stalk eyes and a pair of stout grasping appendages with a terminal spine suggest it was raptorial.Interestingly,the two pear-shaped,three-dimensionally preserved objects that are present in the head region and at the base of the grasping appendages closely resemble the venom glands of some living arthropods in size,shape,and position.These features indicate that the presence of venomous predators could date back 520 million years.Furthermore,our observations suggest that the feeding strategies and organs adapted for this purpose had already reached a high level of diversity and anatomical sophistication in the Early Cambrian ecosystems.     

Earliest record of megaphylls and leafy structures, and their initial diversification

Evolutionary changes in the structure of leaves have had far-reaching effects on the anatomy and physiology of vascular plants, resulting in morphological diversity and species expansion. People have long been interested in the question of the nature of the morphology of early leaves and how they were attained. At least five lineages of euphyllophytes can be recognized among the Early Devonian fossil plants (Pragian age, ca. 410 Ma ago) of South China. Their different leaf precursors or "branch-leaf complexes" are believed to foreshadow true megaphylls with different venation patterns and configurations, indicating that multiple origins of megaphylls had occurred by the Early Devonian, much earlier than has previously been recognized. In addition to megaphylls in euphyllophytes, the laminate leaf-like appendages (sporophylls or bracts) occurred independently in several distantly related Early Devonian plant lineages, probably as a response to ecological factors such as high atmospheric CO 2 concentrations. This is a typical example of convergent evolution in early plants.     

Yunnanozoon是半索动物的远祖

对澄江化石库中Ｙｕｎｎａｎｏｚｏｏｎ（云南虫）重新研究表明，其身体由吻、邻和躯干三部发构成；躯干前段为具有成对鳃孔的咽腔，后段为消化区这种独特的身体造型与现生肠鳃类半索动物几无二致，从鸸 有明这个早寒武世动物是地球上半索动物门中书已知最古老的祖先类型。     

On the Phylum Vetulicolia

1 The Chengjiang Fauna and the four greatest inno- vations in the evolutionary history of animals In 1837, i.e. 22 years before the formal publication of Origin of Species, by comprehending the finch story in the Galapagos, Charles Darwin understood that an animal species could be transformed into another species bygradual modifications and for the first time drew the fa- mous “branching tree”[1]. A century later, passing through two major revisions, i.e. the so-called Neo-Darwinism and …     

High frequency of 'super-cyclones' along the Great Barrier Reef over the past 5,000 years

Understanding long-term variability in the occurrence of tropical cyclones that are of extreme intensity is important for determining their role in ecological disturbances, for predicting present and future community vulnerability and economic loss and for assessing whether changes in the variability of such cyclones are induced by climate change. Our ability to accurately make these assessments has been limited by the short (less than 100 years) instrumented record of cyclone intensity. Here we determine the intensity of prehistoric tropical cyclones over the past 5,000 years from ridges of detrital coral and shell deposited above highest tide and terraces that have been eroded into coarse-grained alluvial fan deposits. These features occur along 1,500 km of the Great Barrier Reef and also the Gulf of Carpentaria, Australia. We infer that the deposits were formed by storms with recurrence intervals of two to three centuries, and we show that the cyclones responsible must have been of extreme intensity (central pressures less than 920 hPa). Our estimate of the frequency of such 'super-cyclones' is an order of magnitude higher than that previously estimated (which was once every several millennia), and is sufficiently high to suggest that the character of rainforests and coral reef communities were probably shaped by these events.     

The earliest-known ancestors of Recent Priapulomorpha from the Early Cambrian Chengjiang Lagerstaette

The taxonomy of an early ancestor of Recent Priapulidae, Xiaoheiqingella peculiaris (= Yunnanpriapulus halteroformis Huang et al., 2004) from the Early Cambrian Chengjiang fossil Lagerstaette, is revised. Morphological characters comprise a pair of caudal appendages rather than a single appendage flanking the trunk end and a possible urogenital duct found inside the preanal region. An additional extremely rare fossil priapulid worm, Paratubiluchus bicaudatus gen. nov., sp. nov. is also described herein. Its diagnostic characters are: an introvert bearing 25 longitudinal rows of scalids, a distinct neck region, no annulus on the oval trunk, and a pair of caudal appendages. The proportion of body parts is similar in size to that of loricate larvae of Recent priapulids and larva-formed Palaeopriapulitidae. Taking account of the features of Xiaoheiqingella, bicaudal appendages are considered to be a synapomorphy of Priapulidae and Tubiluchidae. Paratubiluchus gen. nov. is most likely a candidate for the ancestor of the Tubiluchidae; it probably originated from a larva-formed priapulid with 25 rows of scalids, thus representing an intermediate link between the priapulids in mature-form and the priapulids with lorica.     

鄂尔多斯盆地延长组开阔浅水湖泊风暴沉积

 通过岩心观察,发现鄂尔多斯盆地延长组长8油层组具有典型的风暴沉积特征,主要包括底面侵蚀构造、丘状交错层理、特征异地煤、滞积层、生物逃逸迹等,是风暴流作用于湖岸线附近的三角洲沉积,是沉积物经再搬运和再沉积的产物。其理想的垂向序列由A-G7段构成,向湖盆中心方向,强能量风暴流形成的滞积层和块状砂岩减少,低-弱能量的丘状交错层理和波状纹层段明显增加。鄂尔多斯盆地延长组发育独具特色的开阔浅水湖泊风暴沉积,垂向上表现为多期叠置,平面上表现为对浅水三角洲沉积物的全面改造,总体上呈现出“满盆砂”,成为一种新型储集层,可形成大规模分布的岩性油气藏。     

An armoured Cambrian lobopodian from China with arthropod-like appendages

Cambrian fossil Lagerst?tten preserving soft-bodied organisms have contributed much towards our understanding of metazoan origins. Lobopodians are a particularly interesting group that diversified and flourished in the Cambrian seas. Resembling 'worms with legs', they have long attracted much attention in that they may have given rise to both Onychophora (velvet worms) and Tardigrada (water bears), as well as to arthropods in general. Here we describe Diania cactiformis gen. et sp. nov. as an 'armoured' lobopodian from the Chengjiang fossil Lagerst?tte (Cambrian Stage 3), Yunnan, southwestern China. Although sharing features with other typical lobopodians, it is remarkable for possessing robust and probably sclerotized appendages, with what appear to be articulated elements. In terms of limb morphology it is therefore closer to the arthropod condition, to our knowledge, than any lobopodian recorded until now. Phylogenetic analysis recovers it in a derived position, close to Arthropoda; thus, it seems to belong to a grade of organization close to the point of becoming a true arthropod. Further, D. cactiformis could imply that arthropodization (sclerotization of the limbs) preceded arthrodization (sclerotization of the body). Comparing our fossils with other lobopodian appendage morphologies--see Kerygmachela, Jianshanopodia and Megadictyon--reinforces the hypothesis that the group as a whole is paraphyletic, with different taxa expressing different grades of arthropodization.     

惠民凹陷中央隆起带西部下第三系沙河街组三段上部的风暴重力流沉积

利用岩心观察、粒度分析、薄片研究等方法 ,研究了惠民凹陷中央隆起带西部沙河街组三段上部风暴沉积的主要特征。研究结果表明 ,本区风暴沉积具有①低成分成熟度、高结构成熟度的岩性特征 ;②重力流特征的变形构造 ;③反映风暴浪作用的层理及层面构造 ;④与风浪作用相关的生物成因构造及典型的“似鲍马序列”。本区风暴岩沉积物源来自于湖盆的三角洲及滩坝沉积。按它们之间的相对位置可以划分出两种风暴岩类型 ,即滩前较深水风暴岩和滩后浅水风暴岩。建立了该区的风暴沉积相模式     

惠民凹陷临13断块沙二下储层结构模型

惠民凹陷临１３断块沙二下沉积期主要发育了滨浅湖滩坝和风暴沉积,共划分出６种岩石相类型。储层的储集空间以原生孔隙为主,物性较好,主要受沉积微相或岩相控制,其中滨浅湖滩坝沉积的主体微相或颗粒支撑砂岩相为高孔高渗带。沙二下沉积期的滨浅湖滩坝和风暴沉积相对稳定,非均质性程度低。但是,由于沉积物的粒度偏细,泥质含量高,渗透率的非均质性程度高于孔隙度     

Sponge fossil assemblage from the Early Cambrian Hetang Formation in southern Anhui

Abundant well-preserved large articulated sponge fossils and isolated spicules have been reported from the Early Cambrian Hetang Formation, southern Anhui Province. This unique epifaunal fossil assemblage dominated by articulated sponge fossils is called the Xidi Sponge Fauna. The sponge fauna lived in a quiet oxygenic environment below the storm wave base. Bloom of phytoplankton and rapid sedimentation rate resulted in the deposition of the black shales. Sufficient food supply, lack of other competitors, abundant ecological niches, and demand for oxygen during early Cambrian were in favor of the diversification and evolution of large sponges in the Early Cambrian.     

Stratigraphy and age of the Daohugou Bed in Ningcheng, Inner Mongolia

Recent fieldwork has extended the distribution of the Daohugou Bed deposits from the Daohugou Village to its several neighboring areas. The fossil-bearing Daohugou deposits uncomformably overlie complex bedrocks, and comprise three major parts. The red shales in the lower part were misidentified as belonging to the Tuchengzi Formation. Field excavation has indicated that the shales of upper part of the bed are the major fossil-bearing horizon. Due to strong tectonic activities, sediments were often folded with the sequences inverted in the region. Some newly recognized contacts between the Daohugou Bed and the volcanic rocks showed that the ignimbrite of the Tiaojishan Formation （159-164 Ma） underlies the Daohugou deposits, rather than overlying the latter as previously proposed. Thus, the age of the Daohugou deposits should be younger than the age of the ignimbrite, and thus it was incorrect to correlate the Daohugou Bed with the Middle Jurassic Jiulongshan Formation. Although biostratigraphic studies based on conchostracans and insects support a Middle Jurassic-early Late Jurassic age for the Daohugou deposits, vertebrate fossils such as Liaoxitriton, Jeholopterus and feathered maniraptorans show much resemblance to those of the Yixian Formation. In other words, despite the absence of Lycoptera, a typical fish of the Jehol Biota, the Daohugou vertebrate assemblage is closer to that of the Early Cretaceous Jehol Biota than to any other biota. We propose that the Daohugou fossil assemblage probably represents the earliest evolutionary stage of the Jehol Biota based on both vertebrate biostratigraphy and the sedimentological and volcanic features which suggest the Daohugou deposit belongs to the same cycle of volcanism and sedimentation as the Yixian Formation of the Jehol Group.     

U-series dating of Zhangkou Cave in Yiliang,Yunnan Province:Evidence for human activities in China during 40—100 ka

The cultural deposits at pits T1 and T2 in Zhangkou Cave are intercalated with several flowstone layers. U-series dates show that the capping and 2rid flowstone layers are Holocene of age. The 4th and 5th flowstone layers are ca. 55 and 110 ka old respectively. The lithic artifacts bracketed by them provide unequivocal evidence for hominid presence during this time interval. The ““temporal gap““ of hominid fossil, widely quoted as in support of the out-of-Africa hypothesis, is most probably an artifact due to systematic errors of dating techniques. The infillings under 6th flowstone layer date to ＞300 ka, much older than the previous estimate at 15 ka based on classical 14C dating of fossil bones, providing one more example of the limited reliability of this chronometer. With rich relics and favorable conditions for precise dating, this site is promising for further multidisciplinary studies to address issues concerning recent human evolution in China.