Revision of Aleiodes (Hemigyroneuron) parasitic wasps (Hymenoptera: Braconidae: Rogadinae) with reappraisal of subgeneric limits,descriptions of new species and phylogenetic analysis

The parasitic rogadine braconid wasp subgenus Hemigyroneuron, previously and inappropriately synonymized with Aleiodes (Chelonorhogas) as a result of misidentification, is here synonymized with Aleiodes (Pholichora). A key to the 26 species of Aleiodes (Hemigyroneuron) is presented, including 16 which are described and illustrated as new: A. (H.) amoretae Butcher and Quicke, A. (H.) faddenae Butcher and Quicke, A. (H.) glandularis Butcher and Quicke and A. (H.) plurivena Butcher and Quicke from South Africa; A. (H.) bakeri Butcher and Quicke from Indonesia; A. (H.) bengalensis Butcher and Quicke from India; A. (H.) cusackae Butcher and Quicke from Uganda; A. (H.) elgon Butcher and Quicke, A. (H.) meruensis Butcher and Quicke and A. (H.) pappi Butcher and Quicke from Kenya; A. (H.) fenwickae Butcher and Quicke from Congo; A. (H.) pseudospeciosus Butcher and Quicke from Brunei; A. (H.) roberti Butcher and Quicke from Thailand; A. (H.) sharkeyi Butcher and Quicke from Madagascar; A. (H.) trianguliscleroma Butcher and Quicke from Malawi and Kenya; and A. (H.) ugandaensis Butcher and Quicke from Uganda. A. (H.) apicale (Brues) comb. nov., A. (H.) dubiosus (Fullaway) comb. nov., A. (H.) speciosus (Baker) comb. nov. and A. (H.) suffusus (Baker) comb. nov. are redescribed and illustrated. Males of most species of A. (Hemigyroneuron) are shown to possess a glandular pore opening near the mid-posterior margins of metasomal tergites 4–6, though the situation in A. (H.) dubiosus is different. Results of morphological phylogenetic analysis are presented and indicate clear separation between African and Asian species groups. Two of the new species have been reared from Geometridae and one apparently from Pieridae.     

Metazoan endoparasites of 18 anuran species from the mesophytic semideciduous Atlantic Forest in southern Brazil

Brazil has the richest anuran biodiversity in the world and many species have not yet been studied for parasites. One hundred and forty frogs belonging to 18 species (Rhinella schneideri, Dendropsophus aff. minutus, Dendropsophus nanus, Hypsiboas albopunctatus, Hypsiboas punctatus, Hypsiboas raniceps, Phyllomedusa tetraploidea, Pseudis platensis, Scinax fuscovarius, Trachycephalus typhonius, Leptodactylus chaquensis, Leptodactylus fuscus, Leptodactylus aff. latrans, Leptodactylus mystacinus, Leptodactylus podicipinus, Physalaemus cuvieri, Physalaemus nattereri and Elachistocleis bicolor) collected in aquatic habitats in forest remnants and surrounding agricultural landscapes in southern Brazil were examined for endoparasites. Twenty-four species of endoparasites were identified among 21 adult and three larval individuals: one species of Monogenea, Polystoma lopezromani; eight species of Digenea, Catadiscus marinholutzi, Catadiscus propinquus, Choledocystus simulans, Gordoderina sp., Neohaematoloechus neivai, Rauschiella lenti, Rauschiella linguatula, Rauschiella repandum; one species of Cestoda, Ophiotaenia sp.; one species of Acanthocephala, Centrorhyncus sp.; 12 species of Nematoda, Aplectana sp., Aplectana travassosi, Cosmocerca podicipinus, Raillietnema sp., Schrankiana formolusa, Falcaustra mascula, Oxyascaris oxyascaris, Ochoterenella digiticauda, Physaloptera sp., Brevimulticecum sp., Rhabdias sp., Parapharyngodon hylidae; and one species of Annelida, Dero (Allodero) lutzi. The endoparasite fauna found is constituted by generalist species. Two new records of occurrence in Brazil and 33 new host records are reported.     

Cladistics and taxonomy of the Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera,Empididae)

The Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera: Empididae) is defined on the basis of two synapomorphies, namely epandrial lamellae connected anteriorly and male cerci made up of an anterior almost bare lobe and a posterior bristled lobe. This group includes 10 species: E. (C.) brazzavillensis sp. n., E. (C.) barbitos Smith, E. (C.) juxtaripa sp. n., E. (C.) lyra Smith, E. (C.) plumata sp. n., E. (C.) chrysocera Collin, E. (C.) cuthbertsoni Smith, E. (C.) machipandensis Smith, E. (C.) samaruensis sp. n. and E. (C.) singulare sp. n. All species are described and keyed. The relationships between these species are established and two complexes of species distinguished. The Empis (Coptophlebia) chrysocera-group apparently occurs everywhere in the Afrotropical region except South Africa.     

Copepoda associated with scleractinian corals on the Great Barrier Reef,northeastern Australia,with a key to the genera of the Lichomolgidae

Nine new species and three new genera, Parandrianellus, Scyphuliger, and James-cookina, are associated with various species of hard corals in the vicinity of Lizard Island, Big Broadhurst Reef, and Bowl Reef on the Great Barrier Reef. These copepods include the lichomolgids Parandrianellus annulatus from Hydnophora rigida, Scyphuliger concavipes from Acropora hyacinthus and Acropora sarmentosa, Scyphuliger manifestus from Acropora hyacinthus and Acropora squarrosa, James-cookina redacta from Echinopora horrida, Panjakus eumeces from Hydnophora rigida, Anchimolgus abbreviatus and Anchimolgus tanaus from Acrhelia horrescens, Anchimolgus exsertus from Echinopora horrida, and Odontomolgus bulbalis from Merulina ampliata. Schedomolgus tenuatus Humes, 1990, recorded from Acropora hyacinthus and Acropora humilis, is transferred to the genus Scyphuliger. Schedomolgus lobophorus (Humes and Ho, 1968) is recorded from Acropora hyacinthus, Acropora humilis, Acropora sarmentosa, and Acropora intermedia; Amarda goniastreae Humes, 1985, from Favia stelligera; and Amardopsis merulinae Humes, 1974, from Merulina ampliata and Leptoria phrygia. The harpacticoid Alteuthellopsis corallina Humes, 1981, is reported from Merulina ampliata and Stylophora pistillata. A list of those copepods previously known to be associated with hard corals in the area and a key to the 81 genera of the Lichomolgidae are provided.     

An identification guide to Eurysternine dung beetles (Coleoptera,Scarabaeidae)

An identification guide to the genus Eurysternus is presented. One new species, confusus is described from South America. The synonymies inflexus (Germar) (=planipennis Lucas), parallelus Castelnau (=femoralis Lucas), hirtellus Dalman (=cirratus Harold), calligrammus Dalman (=opacus Lucas), foedus Guérin-Méneville (=claudicans Kirsch), velutinus Bates (=hypocrita Balthasar), and caribaeus (Herbst)(=peruanus Harold) are established, and four previously established synonymies confirmed. Lectotypes are designated for the following species of Eurysternus: mexicanus Harold, plebejus Harold, angustulus Harold, velutinus Bates, hypocrita Balthasar, hamaticollis Balthasar, nebulosus Kirsch, peruanus Harold, sulcifer Balthasar, cirratus Harold, planipennis Lucas, cyanescens Balthasar, femoralis Lucas, and for Onitis deplanatus Germar.

Neotypes are designated for the following species of Eurysternus: magnus Castelnau, marmoreus Castelnau, impressicollis Castelnau, foedus Guérin-Méneville, hirtellus Dalman, opatrinus Perty, parallelus Castelnau, cayennensis Castelnau, and for Scarabaeus caribaeus Herbst.

Type-species are designated for the genera Eurysternus Dalman and Aeschrotes Saint-Fargeau & Serville.     

Classification,reconstructed phylogeny and geographic history of the Neotropical phorid flies of the genus Gymnophora (Diptera: Phoridae)

The Neotropical species of Gymnophora are revised to include 28 species, 17 of which are new. Two subgenera, Cerocratia (in a revised sense) and Gymnophora s.s. are recognized. Cerocratia includes Gymnophora triangularis sp. nov., G. acutangula Schmitz, G. latibrachia sp. nov., G. lacertosa sp. nov., G. chilensis Borgmeier, G. parachilensis sp. nov., G. aemula Borgmeier, G. forticornis Schmitz, and doubtfully G. colona Brues. G. adumbrata Borgmeier is considered as a junior synonym of G. aemula (syn. nov.). Within Gymnophora s.s. there are four major lineages, the G. commotria-group, the G. carina-group, the G. cymatoneura-group and the G. quartomollis-group. Of these, the G. carina-group and the G. quartomollis-group are known only from the Holarctic Region. The G. commotria-group consists of the G. forcipis-series, containing G. forcipis sp. nov., subuncata sp. nov. and uncata sp. nov., and G. damula-series, whose species are further organized into the G. strigula-subseries, containing G. strigula sp. nov. and G. unidentata sp. nov., plus the G. falciformis-subseries, containing G. alces sp. nov., damula sp. nov. and falciformis sp. nov. Also included in the G. commotria-group are eight residual species whose relationships cannot be deduced at this time. Three of these species, G. commotria Schmitz, penai sp. nov. and quadrata sp. nov. are more closely related to each other than to G. auricula sp. nov., brasiliensis Borgmeier, heteroneura Schmitz, inusitata sp. nov. and trispina sp. nov. The G. cymatoneura-group contains G. cymatoneura Enderlein and G. spiracularis Borgmeier, and the male of G. spiracularis is described for the first time. A key is given that permits identification of males of most species, but females cannot be identified with certainty. A partially sequenced classification of the genus indicates the relationships of most of the included species. The genus may have originated in the Neotropical Region, although the evidence is tenuous, and shows some similarity of distribution to Andean-centered plant groups.     

Gastrointestinal helminths of eleven species of Emoia (Squamata: Scincidae) from Papua New Guinea

Four hundred and forty‐six individuals representing 11 species of Emoia (Scincidae), E. atrocostata, E. battersbyi, E. caeruleocauda, E. impar, E. jakati, E. kordoana, E. longicauda, E. obscura, E. pallidiceps, E. physicae and E. popei, collected in Papua New Guinea were examined for helminths. Two species of Cestoda (Adenobrechmos greeri, Cylindrotaenia hickmani); five species of Digenea (Mesocoelium monas, Paradistomoides gregarium, Saurokoilophilia kinsellai, Spelotrema brevicaeca, Zeylanurotrema sphenomorphi); 14 species of Nematoda, (gravid individuals of Abbreviata oligopapillata, Hedruris hanleyae, Kreisiella chrysocampa, Maxvachonia adamsoni, Meteterakis crombiei, Parapharyngodon maplestoni, Physaloperoides milnensis, Pseudoreticularia dipsarilis, Raillietnema nanus, Skrjabinodon sp. 1, Skrjabinodon sp. 2 and larvae (in cysts) of Abbreviata sp., Acuariidae gen. sp., and Riticulariidae gen. sp.) were found. The mean helminth species richness for infected skinks was 6.4±4.7 SD (range 1–14 species). Fifty‐three new host records are reported.     

The Stegana shirozui species group (Diptera,Drosophilidae) from East Asia

One subgroup of the Stegana (Steganina) coleoptrata species group is elevated as a species group (i.e., the shirozui species group), so as to include the three known species, S. ctenaria Nishiharu, S. (S.) masanoritodai Okada and Sidorenko and S. (S.) shirozui Okada, as well nine new species reported from China and Japan: S. (S.) jiajinshanensis, S. (S.) jianfenglingensis, S. (S.) maoershanensis, S. (S.) multidentata, S. (S.) qinlingensis, S. (S.) saigusai, S. (S.) shennongi, S. (S.) wuyishanensis and S. (S.) xanthosticta spp. nov. A key to all the species of this group is provided.     

Species of the genera Bakera and Mangganeura (Homoptera: Auchenorrhyncha: Typhlocybinae) occurring on mango and other crop plants in the Oriental region

The genera Bakera (Bakera), Bakera (Guinobata) and Mangganeura (Homoptera: Auchenorrhyncha: Typhlocybinae) are re-described and compared. Sandalla is considered a synonym of Bakera (Guinobata) (syn. nov.).

Bakera (Bakera) nigrobilineata (Melichar) (=vittata Mahmood) is redescribed and illustrated from mango in the Philippines, on which it is frequently abundant, and found in association with Bakera (Bakera) luzonica sp. nov. which is described. Bakera (Bakera) abaca sp. nov. is described from the Philippines having been collected on abaca (Manila hemp). Bakera (Guinobata) nigroscuta Mahmood is redescribed from the Philippines and its association with coconut confirmed. Mangganeura reticulata Ghauri, described from Malaysia on mango, has been found in Thailand and the hitherto unknown female is described.     

A revision of East Asian Acanthomysis (Crustacea: Mysida: Mysidae) and redefinition of Orientomysis,with description of a new species

Twenty species of Acanthomysis s. l., distributed in Japanese, Korean and Chinese coastal waters, are revised. These species are distinguished from Acanthomysis s. str. by the following characters: the carpopropodus of the endopod of the third to eighth thoracic limbs divided into three to eight subsegments; the exopod of the fourth male pleopod with two long subequal terminal setae; the telson long linguiform or triangular without basal dilated portion and armed with spines throughout margins. On the basis of these characters, the genus Orientomysis, which was revived by Holmquist () and comprised O. japonica and O. mitsukurii, is redefined and recognized to contain the following 18 valid species: O. aokii, O. aspera, O. crassispinosa, O. fujinagai, O. hwanhaiensis, O. koreana, O. leptura, O. meridionalis, O. okayamaensis, O. pseudomitsukurii, O. robusta, O. rotundicauda, O. sagamiensis, O. serrata, O. sheni, O. tamurai, O. tenella, and O. tenuicauda. Acanthomysis longicauda and A. nakazatoi are synonymized with O. rotundicauda and O. japonica, respectively. A new species, O. arenaria, is described from Japan.     

The flower bug genus Orius Wolff, 1811 (Hemiptera: Heteroptera: Anthocoridae: Oriini) of Thailand

The flower bug genus Orius Wolff, 1811 (Hemiptera: Heteroptera: Anthocoridae: Oriini) in Thailand is reviewed. Eleven valid species are recognised; seven of them are described as new to science: Orius (O.) sakaerat, O. (O.) taksini, O. (O.) tomokunii, O. (O.) filiferus, O. (O.) machaerus, O. (O.) inthanonus and O. (Trichorius) crassus. Orius (Heterorius) dravidiensis Muraleedharan, 1977, which has been known from India, is recorded from Thailand for the first time, and is correctly placed in the subgenus Dimorphella Reuter, 1884. The subgenus Paraorius Yasunaga and Miyamoto, 1993 is proposed as a synonym of Dimorphella. Diagnoses, digital habitus images, scanning electron micrographs and illustrations of diagnostic features including both male and female genitalia are provided. Keys to the Thai species are offered to facilitate identification. Biology of Thai species is also discussed.     

Nine new species of the subgenus Stegana (Oxyphortica) from the Oriental region,with morphological and molecular evidence based on Chinese species (Diptera: Drosophilidae)

Nine new species of the subgenus Stegana (Oxyphortica) have been discovered from the Oriental Region: S. (O.) chuanjiangi Zhang and Chen, sp. nov., S (O.) dainuo Wang and Chen, sp. nov., S. (O.) dawa Zhang and Chen, sp. nov., S. (O.) laohlie Zhang and Chen, sp. nov., S. (O.) luchun Wang and Chen, sp. nov., S. (O.) mengwan Wang and Chen, sp. nov., S. (O.) triodonta Cheng and Chen, sp. nov. from southern China; S. (O.) dehiscens Cheng and Chen, sp. nov. and S. (O.) spinosa Cheng and Chen, sp. nov. from eastern Malaysia. One newly recorded species from China: S. (O.) maichouensis Sidorenko is re-described. The DNA sequences of the mitochondrial COI gene with BOLD Process ID and GenBank accession numbers are provided for the above seven new species and five known species from China: S. (O.) acutipenis Xu, Gao and Chen, S. (O.) adentata Toda and Peng, S. (O.) hirtipenis Xu, Gao and Chen, S. (O.) latipenis Xu, Gao and Chen, and S. (O.) maichouensis Sidorenko. Intra- and interspecific pairwise K-2P distances among these species are summarized and the phylogenetic relationships are reconstructed based on the neighbour-joining (NJ) analyses.

www.zoobank.org/urn:lsid:zoobank.org:pub:2FA403F6-5535-40A4-9FD0-ADB9913EDE83.     

Seven new species of Amynthas (Clitellata: Megascolecidae) and new earthworm records from Taiwan

Earthworm specimens collected in southern Taiwan consisted of seven new species of Amynthas and several previously known species, mostly widely distributed peregrines. The new species are A. nanrenensis of the octothecal A. corticis species group, A. monsoonus and A. huangi of the sexthecal A. aelianus species group, and four proandric octothecal species: A. chaishanensis, A. hengchunensis, A. kaopingensis and A. ailiaoensis. Amynthas chaishanensis has dorsal intrasegmental spermathecal pores, but the other three proandric species have dorsal, lateral or ventral intersegmental spermathecal pores, respectively. The proandric species are united by several features, including the enclosure of segment xi in a sac, as in A. formosae (also proandric), octothecal with spermathecae in vi–ix, spermathecal diverticula stalks generally kinked and often enclosed in membrane, and prostatic ducts divided polytomously into numerous small ductlets, which may be grouped into bundles of two to five. In A. ailiaoensis the prostatic duct trunk contains up to seven separate lumens in the ental half, surrounded by the circular muscle of the duct, while in A. chaishanensis the prostatic duct trunk contains about 40 small lumens. Of the previously known species in the collection, Pontodrilus litoralis and Metaphire houlleti are first reported from Taiwan. Additional locations for A. incongruus and A. robustus are given, and in the latter case the material appears to be the usual male‐sterile morph. Other species found are Pontoscolex corethrurus, Amynthas corticis, A. gracilis, Metaphire californica, and Polypheretima elongata.     

Taxonomy and cladistic analysis of the subgenus Xenomorellia Malloch (Diptera: Muscidae: Morellia Robineau-Desvoidy) with description of two new species

Xenomorellia Malloch, a subgenus of Morellia Robineau-Desvoidy, is revised to include two new species, Morellia (Xenomorellia) inca Nihei and Carvalho sp. nov. from South America, and M. (X.) maia Carvalho and Nihei sp. nov. from Costa Rica and Mexico. Diagnoses for M. (X.) holti (Malloch) and M. (X.) montanhesa (Albuquerque) are provided, as well as an identification key to the four species of the subgenus. A cladistic analysis was performed to test the monophyly of Xenomorellia and to recover the phylogenetic relationships among its species. Tree searches resulted in one single most-parsimonious cladogram, wherein the monophyly of Xenomorellia is supported, as well as a sister-group relationship with the Neotropical subgenus Trichomorellia Stein. Xenomorellia was divided into two clades: one with Caribbean–Andean species (maia?+?inca), and another with species from southeastern South America (holti?+?montanhesa).     

Description of new Ceratitis MacLeay (Diptera,Tephritidae) species from Africa

As an addition to earlier published systematic revisions of the genus Ceratitis MacLeay, five new species are hereby described: Ceratitis (Pardalaspis) millicentae sp. n., C. (Ceratalaspis) oraria sp. n., C. (C.) perisae sp. n., and C. (C.) perseus, sp. n. all from Kenya, and C. (C.) ealensis sp. n. from the Democratic Republic of Congo. Their relationship with closely allied species within the respective subgenera is discussed and differentiating characters given.     

Smicridea McLachlan (Trichoptera: Hydropsychidae) from northwestern Argentina and Bolivia: new species,redescription, association and new records

Four new species of Smicridea (Rhyacophylax) are described from Argentina and Bolivia. Smicridea (Rhyacophylax) bifida sp. nov., S. (R.) elisae sp. nov. and S. (R.) valeni sp. nov. are described from the male; S. (R.) thermophyla sp. nov. is described as the first species of Smicridea from thermal springs. The larva and pupa of S. (R.) thermophila were associated using the metamorphotype method. New distributional records of S. (R.) dithyra, S. (R.) atrobasis, S. (R.) peruana and S. (R.) pampeana are included. Morphological characters of male S. (R.) peruana that were not mentioned before are included. Illustrations of all taxa named in this paper are included.     

Grass-breeding fruit flies and their allies of Africa and Asia (Diptera: Tephritidae: Ceratitidinae)

The African genera Bistrispinaria Speiser, Clinotaenia Bezzi and Leucotaeniella Bezzi are reviewed. Clinotaenia camerunica sp. nov. and Leucotaeniella mambillae sp. nov. are described. Clinotaenia atlas Munro is transferred to Bistrispinaria Speiser and Carpophthoromyia superba Bezzi is transferred to Clinotaenia, whilst B. uranos (Hering) is placed as a new synonym of B. woodi (Bezzi) and C. cedarensis Munro is placed as a new synonym of C. grata (Wiedemann). Notes are included on the Asian genera Anoplomus Bezzi, Proanoplomus Shiraki, Rhaibophleps Hardy and Sinanoplomus Zia, all of which appear to be related to Clinotaenia and Leucotaeniella, with Sinanoplomus fasciatus (Walker) transferred from Gastrozona Bezzi. Genus Ceratitoides Hendel is also discussed, with Pardalaspinus Hering, Notophosa Zia and Ceratitisoma Zia placed as new synonyms. The African Chelyophora frigida Hering is transferred from Bistrispinaria and placed as a new synonym of Ceratitoides nigromaculatus Hendel. The Asian C. aberratus (Hardy), C. adnatus (Hardy) and C. maai (Chen) are transferred from Acroceratitis Hendel. Ceratitoides namtamai sp. nov. and C. sikhimensis sp. nov. are described from SE Asia. Keys to all known species are provided. Hostplants are known only for Bistrispinaria, which breeds in various grasses, including guinea grass, sorghum and maize, belonging to the subfamily Panicoideae.     

New species and new records of Veliidae from Espírito Santo State and adjacent Minas Gerais State,Brazil, with notes on nomenclature (Insecta: Heteroptera: Gerromorpha)

A review of the Veliidae occurring in Espírito Santo State, Brazil, is presented. The following new taxa are described: Oiovelia brasiliensis sp. nov., Paravelia capixaba sp. nov., Rhagovelia vaniniae sp. nov., R. mangle sp. nov., R. teresa sp. nov., R. denticulata sp. nov. and R. sooretama sp. nov. Husseyella halophila, Microvelia ayacuchana, and Stridulivelia quadrispinosa are recorded for the first time from southeastern Brazil. Microvelia longipes, M. mimula, M. venustatis, R. accedens, R. bocaina, R. elegans, R. hambletoni, R. itatiaiana, R. robusta, R. tenuipes, R. triangula, R. trianguloides, R. zela and S. tersa are recorded for the first time from Espírito Santo State. The occurrence of H. diffidens in Espírito Santo is confirmed. Additional records from the Caparaó mountain range are presented for M. mimula, R. accedens, R. aiuruoca and R. triangula. A necessary nomenclatural note concerning R. ochroischion, R. sbolos, R. trianguloides and R. turmalis is presented.     

New species and records of Flabellifera from the Indian Ocean (Crustacea: Peracarida: Isopoda)

In this study of flabelliferan Isopoda from the Indian Ocean, 19 species are described as new, in 11 genera and two families: family Cirolanidae, Baharilana koloura, B. lira, Cirolana aldabrensis, C. mimulus, C. somalia, C. undata, Eurydice paxilli, and Metacirolana chemola; family Sphaeromatidae, Cassidinidea clarkae, Cymodoce fuscina, C. lirella, Dynamenella alveolata, D. remex, Heterodina (new genus), H. mccaini, Oxinasphaera brucei, O. furcata, O. tetrodon, Paracilicaea stauros, and Sphaeromopsis sulcifera. Diagnoses of the genera and species are provided, and keys to the Indian Ocean species of most of the genera are provided. The material comes primarily from Aldabra Atoll, the granitic Seychelles islands, Zanzibar Island, and Phuket in Thailand. A few miscellaneous samples from the South African Museum collections, and from the International Indian Ocean Expedition carried out in the mid‐1960s are included.