A review of the north-western Pacific species of the genus Paguristes (Decapoda: Anomura: Diogenidae), I. Five species initially reported by Ortmann (1892) from Japan

Five species of the hermit crab genus Paguristes Dana, 1851, initially reported by Ortmann (1892) from Japan, are revised based on the materials reported by Ortmann (1892), Doflein (1902), Balss (1913) and Miyake (1978), and on newly collected specimens. The specific status of P. palythophilus Ortmann, 1892, is clarified, and a species confused with P. palythophilus is described as new, P. albimaculatus. It has been revealed that P. kagoshimensis Ortmann, 1892, is actually a junior subjective synonym of P. digitalis Stimpson, 1858. The species heretofore assigned to P. kagoshimensis is described as new, P. versus. It has been confirmed that Paguristes acanthomerus Ortmann, 1892, is a valid species and that P. barbatus sensu Ortmann, 1892, is the same species as P. ortmanni Miyake, 1978. Further, a new species, P. doederleini, is described for P. setosus sensu Ortmann, 1892. It is also shown that a part of the group of specimens identified as Paguristes puniceus by Miyake (1978) is referable to P. doederleini. In addition to the three new species recognized herein, the four previously described species are also redescribed in detail in order to make clear their taxonomic status.     

On the correct name for Hemigrapsus edwardsii (Hilgendorf, 1882) (Brachyura: Varunidae) from New Zealand

The specific name for the New Zealand crab long known as Hemigrapsus edwardsii (Hilgendorf, 1882) has been uncertain because of problems with generic definitions. Resolving this question required examination of the relationships between Brachynotus de Haan, 1835, Heterograpsus Lucas 1849 and Hemigrapsus Dana, 1851. The genus Heterograpsus is an objective junior synonym of Brachynotus since they both have the same type species, Gonoplax sexdentatus Risso, 1827. Comparison of Brachynotus and Hemigrapsus shows that the New Zealand species belongs in Hemigrapsus and should be known as H. sexdentatus (H. Milne Edwards, 1837).     

A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae)

The genus Pandalus Leach, 1814, is revised based upon the abundant material from collections in various museums or institutions in the world. Altogether 20 species are recognized which appear to form five groups. Nineteen species are recognized in Pandalus, three of which are described as new: P. curvatus from southern Japan, and P. chani and P. formosanus from Taiwan. Four informal species groups are also recognized within the genus: P. montagui group (P. montagui Leach, 1814; P. borealis Krøyer, 1838; P. goniuruStimpson, 1860; P. jordani Rathbun, 1902; P. tridens Rathbun, 1902; and P. eous Makarov, 1935); P. stenolepis group (P. stenolepis Rathbun, 1902; P. curvatus sp. nov.); P. hypsinotus group (P. hypsinotus Brandt, 1851; P. danae Stimpson, 1857; P. prensor Stimpson, 1860; P. gracilis Stimpson, 1860; P. gurneyi Stimpson, 1871; P. nipponensis Yokoya, 1933; P. teraoi Kubo, 1937; P. chani sp. nov.; and P. formosanus sp. nov.); and P. platyceros group (P. platyceros Brandt, 1851; and P. latirostris Rathbun, 1902). The P. platyceros group appears to be most closely related to the genus Pandalopsis Bate, 1888, but Pandalus is retained as a possible paraphyletic group. Protandrous hermaphroditism is known in all but two species of the genus (P. curvatus and P. formosanus), for which only specimens of either male or female have been available. Pandalus propinqvus G. O. Sars, 1870, is transferred to a new monotypic genus Atlantopandalus, because of its lack of hermaphroditism and possession of some unique morphological characters, including one indicating a close relationsip to Dichelopandalus Caullery, 1896. All species are fully described and illustrated. The affinities and important morphological variations of the species are discussed. A key for adults is presented for the identification of the species. Biogeography of the genus is briefly discussed.     

Studies on the provenzanoi and other pagurid groups: VI. Larval and early juvenile stages of Pagurus ochotensis Brandt (Decapoda; Anomura; Paguridae) from a northeastern Pacific population,reared under laboratory conditions

Zoeal, megalopal, and early juvenile stages of Pagurus ochotensis (Brandt, 1851) from a northeastern Pacific population are described from laboratory-reared individuals. Zoeal and megalopal development in this population is compared with that described for a population of this species from Hokkaido, Japan. First and second juvenile crab stages of this species are described for the first time. The nearly identical developmental morphology of the two populations supports the earlier synonymy of the northeastern Pacific species, Pagurus alaskensis (Benedict) with P. ochotensis.     

North Pacific species of the amphipod (Crustacea) family Stegocephalidae,with description of one new species and redescription of another

The genera and species of the family Stegocephalidae Dana, 1852 (Crustacea: Amphipoda) of the North Pacific, including the Bering Sea, are surveyed, and a key to the species is presented. One new species, Stegocephalus beringi n. sp., is described, and Stegocephalopsis ampulla (Phipps, 1774) is redescribed; S. vegae Oldevig, 1959 is synonymized with S. ampulla. Comments are given on the morphology of three additional species.     

Larval development and emigration behaviour during sea-to-land transition of the land hermit crab Coenobita brevimanus Dana, 1852 (Crustacea: Decapoda: Anomura: Coenobitidae) under laboratory conditions

To determine the early life history of the land hermit crab Coenobita brevimanus Dana, 1852, larvae were cultured individually in the laboratory. The zoeal and the megalopal stages are described and illustrated. The larvae developed through four planktonic zoeal stages to the megalopal stage. The major differences in the zoeal characters between C. brevimanus and other described Coenobita species were found in the armature of the pleomeres, whereas the character of pleomeres of C. brevimanus zoeae is the same as that of the coconut crab Birgus latro, a different genus in the same family. Morphological similarity was also found in segmentations of antennules and antennae in megalopae between C. brevimanus and the coconut crab. Megalopae of C. brevimanus were cultured in containers holding seawater and a hard substrate. These crabs migrate from the sea to land after developing a habit of acquiring gastropod shells.     

Revision of the southern African genus Frankenbergerius Balthasar with description of new taxa (Coleoptera: Scarabaeidae: Scarabaeinae)

The southern African genus Frankenbergerius Balthasar is revised. Two new species, F. opacus sp. n. and F. nitidus sp. n. (South Africa, Western Cape), and one new subspecies, F. armatus tuberculatus ssp. n. (South Africa, Mpumalanga), are described. F. imitativus (Péringuey) is considered a junior synonym of F. forcipatus (Harold). A key to the species and notes on biology are given.     

The genus Monacha in the Western Caucasus (Gastropoda: Hygromiidae)

Seven Monacha species have been found in the Western Caucasus region. Six of these species are endemic there. They all belong to the subgenera Paratheba and Metatheba. Three species are new to science. M. (Paratheba) claussi, sp. n. and M. (Metatheba) subcarthusiana (Lindholm) are the only Monacha species, in which the right ommatophoral retractor sometimes crosses with the penis. This might be due to an ancestral polymorphism, atavism or a basal position of the two species within Monacha, which would mean that Paratheba is paraphyletic and Metatheba is polyphyletic. Monacha (Monacha) cartusiana (O. F. Muller), the only Monacha sensu stricto species in the Caucasus region, has probably been introduced by man. The shell and the genitalia of every species are described and illustrated. The synonyms and all Caucasian locality records are listed. The distributions of the species are illustrated on UTM-grid maps.     

CVIII.—The African spiders of the family Ctenidae in the collections of the British Museum (Natural History)

Summary

The African Ctenidae in the Collections of the British Museum (Natural History) are revised. Thirty species are contained in the subfamilies Cteninae and Acantheinae. Ctenus batesii Pocock, 1903, is a synonym of Ctenus capulinus (Karsch), 1879, Ctenus dirus Thorell, 1899, is a synonym of Ctenus kingsleyi F. O. P.-Cambridge, 1898, Ctenus marshalli F. O. P.-Cambridge, 1898, is a synonym of Ctenus melanogastra (Bösenberg & Lenz), 1895, and Ctenus spenceri F. O. P.-Cambridge, 1898, is a synonym of Ctenus pulchriventris Simon, 1896.

The subfamily Acantheinae is recorded from Africa for the first time. A new genus, Africactenus, is erected for a group of allied species. The genotype is Ctenus agilior Pocock, 1899, and the following new species are described:—A. depressus, A. desartsi, A kribiensis, A. pococki, A. sandersoni, A. simoni, A. sladeni, A. tridentatus and A. trilateralis. Ctenus longurio Simon, 1909, is transferred to the genus Africactenus on examination of the type-material.

Certain species from the Simon Collection are listed in the Appendix and figured for the first time.     

A new fecampiid of the genus Kronborgia (Platyhelminthes: Turbellaria: Neorhabdocoela) parasitic in the intertidal isopod Exosphaeroma obtusum (Dana) from New Zealand

Kronborgia isopodicola sp. n. (Turbellaria: Neorhabdocoela: Fecampiidae) is described from the South Island of New Zealand. The species has a parasitic phase within the haemocoel of the intertidal isopod Exosphaeroma obtusum (Dana). Sexes are separate: each female worm, emerging when mature from its host, secretes a cocoon around itself in which it is joined by a male. Cocoons are found under rocks, are approximately 10 mm long and reflect the shape of the contained female. The worms spawn and lay numerous egg capsules, each containing two embryos. Although placed in Kronborgia Christensen and Kanneworff, 1964 because it is dioecious, K. isopodicola exhibits some characters typical of the genus Fecampia Giard, 1886. It differs from all other described fecampiids by its possession of eyes throughout life.     

Taxonomic observations on the genera Xenocheira Haswell and Ericthonius Milne Edwards (Crustacea: Amphipoda) from Australian coastal waters

The type-species of the genus Xenocheira Haswell, X. fasciata Haswell, is redescribed (including the true male) from Tasmanian material. Pirlot's putative male X. fasciata from Indonesia is allocated to a new species, X. pirloti nom. nov. and K. H. Barnard's ‘X. fasciata’ from the Great Barrier Reef referred to Grandidierella gilesi Chilton. Another Xenocheira species from Western Australia is identified tentatively as X.? seurati Chevreux. Two new species of Ericthonius Milne Edwards are described, one from Tasmania (E. tacitus sp. nov.) and one from Western Australia (E. coxacanthus sp. nov.). The first record of E. pugnax Dana from mainland Australia is claimed for material from Western Australia. The taxonomic relationships of these species are evaluated and discussed.     

The male of Lecane bulla (Gosse, 1851): new support for the synonymy of Lecane Nitzsch,Monostyla Ehrenberg and Hemimonostyla Bartos

The hitherto unknown male of Lecane bulla (Gosse, 1851) is described and figured. It is generally similar to the few known males of other Lecane species. However, whereas the degree of fusion of the toes accords between male and female in other species, this is not the case here: the male of L. bulla has two separate and movable toes, these are completely fused into a single toe in the female. This observation further questions the validity of the taxonomic separation of Lecane in three taxa of generic rank, based on the degree of fusion of the toes. The behaviour and reproduction of Lecane is commented upon.     

Revision of the Traumatomutilla juvenilis species group (Hymenoptera,Mutillidae)

ABSTRACT

Traumatomutilla duplicata duplicata (Gerstaecker, 1874), T. duplicata feia Casal, 1969, T. bruchi André, 1908, T. cristata (Gerstaecker, 1874), T. aterrima (Gerstaecker, 1874) and T. lorena (Cresson, 1902) are proposed as junior synonyms of Traumatomutilla bivittata (Gerstaecker, 1874). Traumatomutilla immaculiceps André, 1901, T. bivittata rubroguttata André, 1901 and T. estrella (Cresson, 1902) are proposed as junior synonyms of T. juvenilis (Gerstaecker, 1874). Traumatomutilla bispiculata (André, 1907) is proposed as a junior synonym of T. miniata (Gerstaecker, 1874). The previously unknown males of T. guarata Casal, 1969 and T. juvenilis are described and illustrated. All species of the T. juvenilis species group are redescribed and illustrated. A new species, Traumatomutilla juvenindica Bartholomay & Williams sp. nov., is described based on males and females from northern South America. Additionally, identification keys to the species and known colour forms of the T. juvenilis species group are provided.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:3563B9EE-A45B-4939-8436-7A808D4F8996     

Palaearctic species of Ammoplanus (Hymenoptera: Sphecidae)

The Palaearctic species of the genus Ammoplanus Giraud are revised, based on material of over 1750 specimens from Europe, North Africa and Central Asia. For almost all species it was possible to examine the type material and in six species the lectotypes have been designated, in two cases (A. perrisi and A. marathroicus) the neotypes. As a result 38 species are recognized as valid, keyed out and commented on. Eight species are described as new and four species names are here placed in synonymy. Several species are of wide distribution, which made it necessary to include also those described from Central Asia and Mongolia. The keys are made to all Palaearctic species but no material is known from the regions east of Mongolia, although four undescribed species are available from South-East Asia. The Afrotropical species are listed and most of them were also examined. Two subgenera are recognized in the region; 32 species are classified in Ammoplanus sensu stricto and six in the subgenus Ammoplanellus. In the nominal subgenus the following species are described as new: A. atlasensis from Morocco, A. biscopula from Israel, A. biskrensis from Algeria, A. denesi from Turkey (Anatolia), A. freidbergi and A. kaplanae from Israel, A. minutus from Bulgaria and Turkey and A. strumae from Bulgaria and Greece. New synonymy: A. angelae becomes a junior synonym of rhodesianus (known before only from Africa), A. crudelis a synonym of A. kaszabi, A. handlirschi a synonym of A. marathroicus, A. suarezi a synonym of A. kohlii, A. subcompressus of A. gegen, A. orbiculatus of A. simplex and A. zarcoi of A. kohlii; A. wesmaeli is confirmed as a synonym of A. perrisi. Otherwise the synonymy is corrected under A. bischoffi and the names A. dusmeti and A. curvidens are re-established as valid. In a short review of the African species A. (Ammoplanellus) cradockensis Arnold, 1959 is placed as a new synonym under A. (Ammoplanellus) latiscapus Leclercq, 1959. The nomenclatural changes are explained under the relevant species.     

Description of Torymus lasallei,sp. nov. (Hymenoptera: Torymidae), a species with an unusual ovipositor

ABSTRACT

Torymus lasallei, sp. nov., a species with an unusual ovipositor, reared from galls of Amphibolips spp. (Hymenoptera: Cynipidae) on Quercus spp. (Lobatae section) from Mexico, is described. The placement of this new species within the genus Torymus, and the morphology and function of its ovipositor, are discussed.

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A review of the genus Parapenaeon Richardson, 1904 (Crustacea: Isopoda: Bopyridae: Orbioninae), with description of three new species from China

Five species of the bopyrid genus Parapenaeon are reported from China, of which three are new to science, and one, Parapenaeon expansa Bourdon, 1979, is a new record. New species are Parapenaeon calculosa, sp. nov., Parapenaeon diatropa, sp. nov. and Parapenaeon sicyoniae, sp. nov., from the East and South China Seas. The female of each new species has six pleomeres, no lateral plates on the sixth pleomere, and biramous uropods, characters that place them in the genus Parapenaeon Richardson, 1904. Each can be distinguished from previously described species of the genus Parapenaeon in body shape, by the characters of oostegite 1, the barbula, brood pouch, pleopods and uropods of the females. A table including all of the currently recognized species of Parapenaeon with type localities, known geographical distribution and reported hosts, is given. A key to the recorded species of Parapenaeon Richardson, 1904 is provided, as is a new diagnosis of the genus.

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