Two new taxa from the Lesser Sundas in the genus Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae)

Two new Indonesian taxa are proposed in Onthophagus Latreille, 1802, illustrating the composite faunal spectrum of the Lesser Sundas (Indonesia): O. montishannoniae, a new species from Flores, and O. accedens nusatenggaricus, a new subspecies from Lombok and Flores. O. accedens Lansberge, 1883 is, with O. dux Sharp, 1875, placed in what is here defined as the dux subgroup within the subgenus Parascatonomus Paulian, 1932. The three included species‐group taxa are keyed, their status is discussed, and new records are given. The position of O. montishannoniae seems taxonomically and geographically isolated: this species has no known close relatives in the region, neither in the Lesser Sundas, nor anywhere else in Wallacea.     

Ooencyrtus marcelloi sp. nov. (Hymenoptera: Encyrtidae), an egg parasitoid of Heliconiini (Lepidoptera: Nymphalidae: Heliconiinae) on passion vines (Malpighiales: Passifloraceae) in Central America

A new species belonging to the genus Ooencyrtus Ashmead is described. Ooencyrtus marcelloi sp. nov. has been reared from eggs of Heliconiini (Lepidoptera: Nymphalidae, Heliconiinae) collected in Panama on Passiflora spp. The new species is compared with its closest Ooencyrtus species, i.e. O. caligo Noyes, O. neustriae Mercet, O. flavipes (Timberlake), O. camerounensis (Risbec), O. endymion Huang and Noyes and Ooencyrtus sp. “undet. C.” (from India). This represents the second record of Ooencyrtus from Heliconinae and the first record of this genus from Panama.     

A revision of the genus Ophiambix (Echinodermata: Ophiuroidea) including the description of a new species

The genus Ophiambix is revised and problems at higher taxonomic levels are discussed. The species Ophiambix aculeatus, O. meteoris and O. devaneyi are redescribed (based on new characters), and a new species O. epicopus is described from the Kermadec Islands, South Pacific. A pictorial key is presented to aid identification.     

Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)

The South American Trogidae are revised. Three genera, Trox Fabricius, Omorgus Erichson (with two subgenera, Omorgus and Haroldomorgus Scholtz), and Polynoncus Burmeister, and 47 species are recorded from the continent and its islands. Trox is represented by the ubiquitous introduced species T. scaber (L.), Omorgus is represented by 13 species and Polynoncus by 33 species. Five new species are described and all others are re-described. A key to all taxa is provided. Adults of all species and their male genitalia are illustrated and their distributions mapped. Observations are presented on phylogeny, zoogeography and biology. The new species are O. capillaceus, O. indigenus, O. nocheles, P. erugatus and P. gibberosus. Four new synonymies are proposed (synonyms listed first): O. triestinae Pittino = P. suberosus (Fabricius); P. furcifer Pittino = P. bifurcatus (Vaurie); P. parafurcatus occidentalis Pittino and P. furcillifer Pittino = P. parafurcatus Pittino. Omorgus badeni (Harold) is reinstated as a valid species. Lectotypes are designated for O. batesi (Harold) and O. badeni (Harold).     

II.—A synoptic revision of the Tanyrrhynchinæ (Curculionidæ)

This paper describes two new earthworm species, Iridodrilus codonothecoformis sp. n. and Iridodrilus condylopapillaris sp. n. and illustrates the major diagnostic features of, and the differences between, Iridodrilus roseus Beddard, 1897 and Heliodrilus lagosensis Beddard, 1890. Hitherto unnoticed features of I. roseus are also illustrated, namely that it possesses a bursa copulatrix and that two pairs of very tiny penial setae in sacs occasionally occur which penetrate the bursa copulatrix. It is also demonstrated that the structures hitherto regarded as a ‘coelomic duct’ that supposedly links the two fertilization chambers is in fact a pair of ovarian ducts that lead from the paired ovaries to the chambers; these ducts are not interlinked. It is further argued that neither Iridodrilus roseus nor Heliodrilus lagosensis can be regarded as having a spermathecal system that is paired internally. The implications of these findings are discussed in relation to the taxonomy of the two named genera. It is recommended that Iridodrilus and Heliodrilus be regarded as valid genera s.s. Beddard, though with modified diagnoses.     

The species of the parasitic bee genus Osirinus (Hymenoptera,Apidae)

The species in the genus Osirinus Roig-Alsina are revised. Seven species are recognized, including O. ruficrus, new species, from south-eastern Brazil and O. tarsalis, new species, from western Brazil and Peru. Osiris parvicollis Ducke and O. santiagoi Almeida are transferred for the first time to Osirinus. An identification key based mainly on females is provided.     

On Anomalotrema Zhukov, 1957, Pellamyzon Montgomery, 1957, and Opecoelina Manter, 1934 (Digenea: Opecoelidae), with a description of Anomalotrema koiae sp. nov. from North Atlantic waters

In order to clarify the systematic position of some opecoeline digeneans parasitic in scorpaenid and gadid fishes from the northern North Atlantic, a study of the genera Anomalotrema Zhukov, 1957, Pellamyzon Montgomery, 1957, and Opecoelina Manter, 1934, was carried out. Anomalotrema is retained in the subfamily Opecoelinae and A. koiae sp. nov. is described for the worms from scorpaenids and gadids which prompted the investigation. This species is distinguished from the Pacific species, A. putjatini Zhukov, 1957, on morphological and biological grounds. Pellamyzon is transferred to the Plagioporinae. A new subfamily, the Opecoelininae subfam. nov., is erected for Opecoelina, which is considered to contain only two or three valid species, O. scorpaenae Manter, 1934, O. helicoleni Manter, 1934 (possibly a synonym of O. scorpaenae) and O. pacifica Manter, 1940. Both O. theragrae Lloyd, 1938, and O. radifistuli (Acena, 1941) are transferred to the genus Podocotyle Dujardin, 1845, as P. theragrae comb. nov. and P. radifistuli comb. nov. O. pharynmagna Annereaux, 1943, is judged to be a synonym of P. radifistuli, and both O. vixiintestina Oshmarin, 1965, and O. vixigastera Oshmarin, 1965, are considered unrecognizable. The possible evolutionary relationships of the subfamilies within the Opecoelidae are commented on.     

Water mites of the family Hygrobatidae Koch from southern Africa (Acari: Hydrachnidia)

New records are given of the water mite family Hygrobatidae (without the genus Atractides) from southern Africa (South Africa, Botswana, Namibia). In addition to one new subgenus of the genus Hygrobates, namely Inflatibates, four new species are described, namely Hygrobates draconicus, H. laceratoides, H. okavangoensis, and H. inflatipes as well as one new subspecies, namely Hygrobates spathuliferus cederberg. The following synonyms are established: the subgenus Capobates Thor with Hygrobates Koch, Hygrobates sanguineus transvaalensis K. O. Viets with Hygrobates spathuliferus (Lundblad), and Ambiguobates permixtus agnewi K. O. Viets with Ambiguobates permixtus K. Viets. The male of Australiobates pilosus (K. Viets) is described for the first time. The synonymization of Diversibates with Australiobates can now be confirmed. Finally, keys are given for the known genera of southern Africa, as well as for all known species of the genus Hygrobates from southern Africa.     

Recent and fossil Lunulitidae (Bryozoa: Cheilostomata) 4. American and Australian species of Otionella

The nominal, free living lunulitiform genus Otionella has been found to include two major groups of species, each one with a distinctive periancestrular pattern of primary zooids. The first group occurs in the Eocene of North America and includes the type species, O. perforata, and several other forms. The second group occurs in the Late Eocene to Recent of Australasia, and may prove to be generically distinct. Intermediate forms, such as O. parvula from the Argentinian Tertiary, are discussed, and a diverse fauna of 10 species from Australia is described. Although some of these species, like O. cupola, O. exigua and O. nitida, have long been known from Australia. it has only recently been recognized that three New Zealand forms, O. squamosa, O. zelandica and O. proberti, also occur in the Tertiary and/or Recent of Victoria, New South Wales, and Western Australia. In addition, four species are considered to be new. These include O. australis, O. auricula and O. minuta, which are probably closely related to one another, and to O. zelandica. The fourth species, O. browni, has a unique periancestrular budding pattern which resembles that of fossil Lunulites more than any form of ‘Australasian Otionella’. Although, in many ways, O. browni could be assigned to Lunulites sensu lato, it is included here in Otionella.     

Red Sea Phyllidiidae (Mollusca,Nudibranchia), with descriptions of new species

All the known Red Sea nudibranchs of the family Phyllidiidae are described and their taxonomic positions discussed. The genus Fryeria Gray, 1853, is regarded as a junior subjective synonym of Phyllidia Cuvier, 1797, and Reyfria gen. nov. is introduced to describe those species differing from Phyllidia primarily in the position of the anus. Of 12 species described here from the Red Sea, four are new to science. Phyllidia arabica Ehrenberg, 1831, is distinguished from Phyllidia varicosa Lamarck, 1801, and Phyllidia sp. is described as the third species confused with this group. A subspecies of P. ocellata Cuvier, 1804, is described and illustrated, as are three new species: P. melanocera, P. monacha and P. multifaria.     

Ophicardelus (Mollusca,Pulmonata) in eastern Australia: how many taxa?

The common south-eastern Australian estuarine ellobiid snails belonging to Ophicardelus have been considered to be one variable taxon by some authors, and three largely sympatric species by others. This study was undertaken to determine the number of taxa present. Different types of data collected included shell shape, allozyme electrophoresis, shell and radular morphology, anatomy and geographic range. These data sets were then examined for consistent discontinuities, and the results showed that the nominal species could be distinguished by a combination of shell shape and sculpture. Individuals from each nominal species also grouped together using allozyme data, although one taxon (Ophicardelus quoyi) is sometimes paraphyletic based on these data. There are several differences in the anatomy and radular morphology that separate the three taxa. These included marked differences in the penial complex, vagina, and presence or absence of a pallial gland. The three taxa also have different geographic ranges, albeit with most of their ranges in sympatry. Based on these differences, Ophicardelus ornatus, O. quoyi and O. sulcatus should be regarded as valid species.     

New species of Orasema (Hymenoptera: Eucharitidae) from Central and South America

ABSTRACT

Six new species of Orasema are described. Four species are placed into a newly recognised lasallei species group: Orasema lasallei sp. nov. (Costa Rica, Honduras, Mexico, Nicaragua), O. janzeni sp. nov. (Costa Rica), O. peckorum sp. nov. (Brazil) and O. vasquezi sp. nov. (Colombia). One species is placed in the wayqecha group, Orasema masneri sp. nov. (Ecuador). One other species is currently unplaced to species group, O. longinoi sp. nov. (Costa Rica). Orasema lasallei was taken from a nest of Pheidole vorax (Fabr.) and an undescribed species near O. longinoi from a nest of Pheidole dossena Wilson (Myrmicinae). The morphology of the digitate labrum is discussed with respect to other Eucharitidae.     

The species of Symplocodes Hebard (Blattodea: Ectobiidae: Blattellinae) with description of a new species from China

This paper provides a generic diagnosis, a species list and taxonomy for the genus Symplocodes Hebard, 1929. The new species Symplocodes euryloba sp. nov. from China is described and illustrated. Symplocodes tsaii Bey-Bienko, 1958 is regarded as a subspecies of Symplocodes marmorata (Brunner von Wattenwyl, 1893), i.e. S. marmorata tsaii comb. nov., and its differences from S. marmorata marmorata (Brunner von Wattenwyl, 1893) are discussed. Two known species, Symplocodes ridleyi (Shelford, 1912) and Symplocodes manubria Feng et Guo, 1990, are redescribed and illustrated, whereas one new specific synonym, i.e. Symplocodes brachialis, is proposed. A key to species worldwide is provided. The tarsal claws of these four species as well as one species of Chorisoserrata Roth, 1998 are examined and photographed to compare the intergeneric differentiation of claw specialization and to discuss its function.     

XXXVIII.—The cretaceous Echinoid genera Infulaster Desor and Hagenowia Duncan

Summary

The Chalk Echinoids of the genera Infulaster and Hagenowia are redescribed. Three published species of Infulaster are regarded as synomyns of the genotype species. Two new species of Hagenowia are described from the English Senonian. While Infulaster is held to be an Echinocorythid of the subfamily Cardiasterinæ, Hagenowia is shown, by certain important structural features, to belong to the curious family Pourtalesiidæ of Loven. The phylogeny of the forms considered is also discussed.     

XVIII.—Some notes concerning the male of Dexamine thea,Boeck

Osteochilotrema malayae gen. nov. sp. nov. (Osteochilotrematinae subfam. nov.) from the intestine of Osteochilus hasselti in Malaysia is described. It is characterized mainly by the symmetrical extracaecal testes and the structure of the male terminal ducts. Protocladorchis pangasii is reported from Pangasius pangasius, Barbus daruphani, B. tambroides and from Malaysia for the first time. It is redescribed and the generic diagnosis amended. Helostomatis indica is reported from O. hasselti in Malaysia, both new records. The subfamily and generic diagnoses are amended. Orientodiscus jumnai was recovered from new hosts, B. tambroides and P. micronemus, and is new for Malaysia.     

New species and records of Flabellifera from the Indian Ocean (Crustacea: Peracarida: Isopoda)

In this study of flabelliferan Isopoda from the Indian Ocean, 19 species are described as new, in 11 genera and two families: family Cirolanidae, Baharilana koloura, B. lira, Cirolana aldabrensis, C. mimulus, C. somalia, C. undata, Eurydice paxilli, and Metacirolana chemola; family Sphaeromatidae, Cassidinidea clarkae, Cymodoce fuscina, C. lirella, Dynamenella alveolata, D. remex, Heterodina (new genus), H. mccaini, Oxinasphaera brucei, O. furcata, O. tetrodon, Paracilicaea stauros, and Sphaeromopsis sulcifera. Diagnoses of the genera and species are provided, and keys to the Indian Ocean species of most of the genera are provided. The material comes primarily from Aldabra Atoll, the granitic Seychelles islands, Zanzibar Island, and Phuket in Thailand. A few miscellaneous samples from the South African Museum collections, and from the International Indian Ocean Expedition carried out in the mid‐1960s are included.     

Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart,Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species

ABSTRACT

The systematics of the arboreal sesarmid crab genera Labuanium Serène and Soh, 1970 and Scandarma Schubart, Liu and Cuesta, 2003 is revised. Also included is the poorly known genus Namlacium Serène and Soh, 1970, which is morphologically similar to Labuanium and may also be arboreal in habit. The present study restricts Labuanium to its type species, L. politum (De Man, 1887), and congeners are transferred to Scandarma, Geosesarma De Man, 1892, and three new genera established herein: Circulium gen. nov., Shinobium gen. nov. and Mindanium gen. nov. Two new species of Scandarma are described from Papua New Guinea and Madagascar. Sesarma jacquinoti Ortmann, 1894, previously regarded as a synonym of Scandarma gracilipes (H. Milne Edwards, 1853) (comb. nov.), is here shown to be a junior synonym of Sesarmops angustifrons (A. Milne-Edwards, 1869). Labuanium schuetteii (Hess, 1865) is regarded as conspecific with Scandarma gracilipes (H. Milne Edwards, 1853) (comb. nov.) and the lectotype of the latter species (here designated) is also selected as the neotype of the former, making the two names objective synonyms. Material which had been identified as ‘Labuanium finni Alcock, 1900’ by Pretzmann (1984) from the South Andaman Islands is actually the poorly known Geosesarma thelxinoe (De Man, 1908), which is here referred to a new genus, Andamanium gen. nov. Sesarma finni Alcock, 1900 s.s. is herein assigned to Scandarma. Namlacium crepidatum (Calman, 1925), the only species in the genus, is redescribed and figured and compared with Labuanium and Scandarma. Keys to species of Circulium and Scandarma are provided.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:414B8DAA-584F-4070-A355-83B583D0D017     

A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae)

The taxonomy of the African representatives of the wolf spider genus Ocyale is revised. Fundamental taxonomic problems were encountered which resulted in confusion in the literature between Ocyale and Pisaura and thus potential instability of the type genus of the family Pisauridae. For reasons of taxonomic stability, a new type species for Ocyale is proposed here: O. neatalanta. Besides the type species, O. dewinterae n.sp., O. discrepans Roewer, 1960, O. grandis n.sp. and Ocyale guttata Karsch, 1878, are described and an identification key is provided. Ocyale spissa (Lenz, 1895) proved to be a junior synonym of O. guttata. Ocyale maculata (L. Koch, 1875), O. spissides Strand, 1907, O. fera Strand, 1908, O. ingenua (Karsch, 1879) and O. conspersa Karsch, 1879, were found to be either nomina dubia or incertae sedis.     

A new troglobitic meenoplid from a lava tube in Western Samoa (Homoptera Fulgoroidea Meenoplidae)

Recent investigations in lava tubes on Savai'i, Western Samoa, have revealed the existence of a new cave-dwelling meenoplid species, Suva oloimoa sp. nov., from the deep cave zone. Troglomorphies displayed by S. oloimoa sp. nov. include reduction of ocelli, compound eyes, pigmentation and, to a lesser degree, of tegmina and wings. Due to these features and to its ecology, the species is regarded as an obligate cavernicole (troglobite). This is the first record of a troglobitic meenoplid in the Pacific.