I.—Notes on the Natural History of East Finmark

Synopsis

A new species of Argyrophylax, parasitic on the Banana Scab-Moth Nacoleia octasema Meyrick, is described and the Oriental and Australasian species of this genus are reviewed; a key is given for their identification. Six species and a subspecies hitherto placed in other genera are here assigned to Argyrophylax, and seven new combinations are established; two specific names are newly placed in synonymy.     

LXVII.—The classification of the thecidean brachiopods

Abstract

The evolution and classification of thecidean brachiopods is discussed. The shell-structure of the geologically earlier genera is shown to be of pseudopunctate pattern, that of the later genera punctate. The pseudopunctate Rhaetic Bactrynium is not considered a thecidean, and the Tiassic Thecospira is shown to be a pseudopunctate spire-bearer. The new genera Moorellina, Vermiculothecidea and Eolacazella are proposed, and the species Moorellina prima (Rhaetic) and Bifolium lacazelliforme (Cenomanian) are described. The possibility of a neotenous origin of the thecideans from the strophomenoids is examined, and the nonthecidean genus Gwynia discussed in connection with this. The heterochronous thecidiform genera are grouped as the subfamily Thecideinae, and the series of thecidelliniforms from which the thecidiforms evolved as Thecidellininae, the two together forming the family Thecideidae, which is placed in the superfamily Thecideacea to express the divergence of thecidean evolution from strophomenoids.     

A new genus and two new species of Philotarsidae. (Insecta: Psocoptera) from East Africa

Summary

Genus Tarsophilus and its species lineatus and aequabilis (Insecta: Psocoptera: Philotarsidae) are described as new. The specimens, all males, were collected in highland areas of East Africa. The new genus is placed in Philotarsidae rather than Pseudocaeciliidae on characters of hypandrium, phallosome, and lacinial tip. Its position in the subfamily Philotarsinae is justified and it is compared to the other genera of this group on both male and on trans-sexual characters.     

The Mollissima subgroup of fannia Desvoidy,with the description of a new species from Burma and a revised key to species (Diptera: Muscidae)

Synopsis

A comparison of the type-species of the genus Coelomyia Haliday (C. mollissima Haliday) with that of the genus Fannia Desvoidy (F. scalaris Fabricius) shows that these two genera cannot be maintained as equivalent phylogenetie taxa. Coelomyia is synonymized with Fannia and its species placed in Fannia, four new combinations thus being established. The mollissima subgroup is defined and its affinities shown, a new species is described, and a revised key to species is given.     

Re-evaluation of the fossil Phoridae (Diptera)

I re-evaluated the known fossil phorids after re-examining much of the Baltic amber and Dominican amber fauna. Many extant genera reported from Eocene Chinese and Eocene/Oligocene Baltic amber are actually not present; instead there are representatives of relatively primitive stem-group taxa. A new genus and species of fossil phorid, Limulomyia tyche, based on a male and a limuloid female, is described from Baltic amber. Some changes in the taxonomy of amber phorids are proposed: Pseudometopina Disney is synonymized with Metopina Schmitz (new synonymy), Diplonevra meunieri Brues is transferred to the genus Godavaria Brown (new combination), Protophorites fimbriatus Brues is considered a junior subjective synonym of Protoplatyphora tertiaria Brues (new synonymy) and Phalacrotophora tertiara Statz is transferred to the genus Gymnophora (new combination). The oldest confirmed dates for extant phorid genera are given.     

Some new African Ulopini (Homoptera,Cicadellidae, Ulopinae)

Summary

Two new genera and two new species of Ulopini are described from Africa; the distribution, and some aspects of the evolution, of genera in this tribe are discussed. It is suggested that Ulopa damaspina Dlabola belongs to the genus Moonia Distant and a new synonmy is suggested. In an Appendix the genera and species of the Ulopini are listed according to their geographical distribution.     

A new genus with three new species of copepods from the family Diaixidae (Crustacea: Calanoida), and a redefinition of the family

Plankton samples taken in the Gulf of Carpentaria, Australia, yielded material of a new genus containing 3 new species: Anawekia spinosa gen. nov. et sp. nov., A. robusta sp. nov. and A. bilobata sp. nov., which are described herein. The family Diaixidae now contains 12 described species in 2 genera, 9 being in the genus Diaixis. It is noted that 3 of the Diaixis species (including the type species) share some features which are not shared with the remaining 6 species, and that these latter species should probably be placed into a third genus. An amplified redefinition of the family is given.     

Plantes Cryptogames de France. Par I. B. H. I. Desmazières. Fasc. XVII. Lille, 1836

Summary

A new genus and species of pilargid polychaete, Litocorsa stremma, is described from silty grounds on the west coast of Scotland. Its relationship to other pilargid genera is discussed. Notes on two other pilargid species and a revised key to the genera of the Pilargidae are given.     

Stridulatory organs of the elm bark beetles Scolytus multistriatus Marsham and Scolytus scolytus Fabricius

Summary

In Scolytidae, stridulatory organs are not usually found in the sex which selects the breeding site and begins the gallery. Exceptions in the literature refer to the genera Dendroctonus and Scolytus. The presence of stridulatory organs in both sexes of the elm bark beetles S. multistriatus and S. scolytus is reported here. The structure of the stridulatory apparatus is described, and the possible role of stridulation in the behaviour of these species is discussed.     

The rediscovery of Danielssenia intermedia Wells 1965 (Copepoda,Harpacticoida): a missing link between the ‘danielsseniid’ genera and Paranannopus Lang 1936 (Paranannopidae)

Danielssenia intermedia Wells 1965, originally described from a single male specimen from the Fladen ground in the North Sea, is redescribed on the basis of new material of both sexes from off Cullercoats, also in the North Sea. Although D, intermedia has many similarities to the genera Danielssenia Boeck and Psammis Sars, it must be placed in a new genus Fladenia based on distinctive features of the female antennule, the mandible, maxilliped, first and fifth legs and the sexual dimorphism of both rami of legs 2–4. The characteristics of Fladenia are such that it serves to link the other genera of the danielsseniid complex with Paranannopus Lang. The proposal that all these genera should be placed in the family Paranannopidae Por is reinforced by close similarities in the structure of the female genital field.     

The Villa group of genera in Africa and Eurasia with a review of the genera comprising Thyridanthrax sensu Bezzi 1924 (Diptera: Bombyliidae)

Summary

Bombyliidae are particularly abundant in arid regions where the greatest diversity of genera and species is found. The Villa group of genera (Anthracinae, Exoprosopini) are especially well represented in the Saharo-Sindian desert region. Difficulty in segregating genera has resulted from reliance on single key characters. Using a suite of characters, including the spermathecae largely overlooked previously, a key is provided to the eleven known genera in the Old World. The genus Thyridanthrax sensu Bezzi (1924) is reviewed and five genera are recognized: Thyridanthrax Osten-Sacken, Exhyalanthrax Becker, Pachyanthrax François, Veribubo Evenhuis, and Caecanthrax gen. nov. Species are allocated to these genera and new species Pachyanthrax laconae, Veribubo gazella and V. saffra and Caecanthrax auratus are also described.     

Diptera from Nepal. Sepsidae

Summary

The material examined included six species in three genera. No new taxa are described but Parapalaeosepsis Duda and related taxa are discussed. Dicranosepsis Duda is placed in synonymy under Parapalaeosepsis, and Parapalaeosepsis bicolor (Wied.) is proposed as a new combination with Sepsis pubipes Brun. as a new synonym.     

Disjunct distribution of Szeptyckiella gen. nov. from New Caledonia and South China undermines the monophyly of Willowsiini (Collembola: Entomobryidae)

Morphology of scales is an important taxonomical character at all levels of Entomobryidae taxonomy. A new scaled genus of Collembola, Szeptyckiella gen. nov., and three new species are described: Szeptyckiella boulouparica sp. nov. and Szeptyckiella sinelloides sp. nov. from New Caledonia, and Szeptyckiella lii sp. nov. from South China. The genus, assigned to Willowsiini, is characterized by pointed scales on the body, eyes and pigment absent, antennal apical bulb absent, dens lacking spines and scales, and bidentate mucro with a short basal spine. It is closest to Hawinella from Hawaii but the latter possesses a falcate mucro. However, its morphological characteristics and its disjunct distribution raise a number of problems. Its widely disjunct distribution is difficult to explain in terms of palaeobiogeographical or more recent dispersal events. The new genus is similar to Sinella without consideration of scales, although both genera are placed in different tribes following the current supra-generic classification of Entomobryidae. We discuss other cases of paired genera differing only by the presence or absence of scales, and placed in either Willowsiini or Entomobryini. On this basis, we assume that scale presence could independently occur in the former tribe, questioning the monophyly of the tribe Willowsiini.     

Revision of the Palaearctic species of the genus Clinocentrus (Hymenoptera,Braconidae)

Revision of the Palaearctic species of the genus Clinocentrus Haliday is given. The composition and geographic distribution of the genus Clinocentrus and the probable generic position of fossil species described by Brues (1933) as Clinocentrus are discussed. The systematic position of this genus in Exothecinae is also discussed and arguments presented to support the differentiation of exothecines and rogadines. Nine species of Clinocentrus are figured and re-described. Three new species and two new subspecies are figured and described: C. kalmyk from Kalmykia, C. kozlovi from Mongolia, C. rhysipoloides from Primorsk Territory of Russia, C. umbratilis ssp. disruptus from Taiwan and C. vestigator ssp. orientalis from Primorsk Territory of Russia. C. stigmaticus Marshall, 1897 and C. jaroshevskyi Telenga, 1941 are synonymized with C. vestigator (Haliday, 1836); C. tarsalis Ashmead, 1894—with C. exsertor (Nees, 1812) and C. gracilipes (Thomson, 1891)—with C. cunctator (Haliday, 1836), Lectotypes of Exothecus analis Wesmael, 1838 and C. antefurcalis Granger, 1949 are designated. The new combination is given: Rhysipolis antefurcalis (Granger). A key to 12 Palaearctic species of Clinocentrus is given.     

The genus Teuchophorus in South-East Asia and New Guinea,description of new species,species-groups and their phylogeny (Insecta,Diptera, Dolichopodidae)

Forty new Teuchophorus species are described and illustrated from Papua New Guinea, Indonesia and Thailand. New data are given for 14 of the 16 species described from Papua New Guinea by Meuffels and Grootaert in 1986. The genera Mastigomyia Becker, 1924 and Olegonegrobovia Grichanov, 1995 are placed in synonymy with the genus Teuchophorus Loew, 1857. Various species-groups are recognized and described and a phylogenetic relationship is proposed. A key is given for the 63 species occurring in South-East Asia and New Guinea, and a world checklist is added.     

VIII.—Notes on the Silurian Brachiopod genera Delthyris,Uncinulina, and Meristina

Summary

In this paper the different interpretations of the genus Delthyris are discussed, and that given by Schuchert in 1913 is adopted with slight amendment. The species Delthyris elevata, D. crispa, D. elegans (nom. nov. for D. crispa, Hisinger, non Linnæus), and D. tenuilamellata, sp. n., from the British Silurian, are all described and figured.

The distinguishing characters of the genera Uncinulus and Uncinulina are pointed out, and Uncinulina is retained with U. fallaciosa as genotype. Rhynchonella stricklandi and Terebratula lewisi are transferred to the genus Uncinulina. Meristina tumida (Dalman) is shown to be a synonym of Meristina obtusa (Sowerby).

The British Silurian Spirifers can be separated into two groups:—(1) those with longitudinal folds and a transverse ornament of imbricating lamellæ (=Delthyris of Dalman) and (2) those with longitudinal folds bearing fine continuous longitudinal striæ (=Eospirifer of Schuchert).

The species referable to both these genera have as a rule been assigned to the genus Spirifer, from which, however, they differ in internal characters.     

Australian thrips of the Haplothrips lineage (Insecta: Thysanoptera)

Keys are presented to the 11 genera and 50 species, including 21 new species, of Thysanoptera in Australia that are related to the worldwide genus Haplothrips Amyot and Serville. These taxa belong to what, in recent literature, has been called the “Haplothrips‐lineage”, that is, one of the three major radiations among the 2700 species and 350 genera of Thysanoptera Phlaeothripinae. The group is redefined, and the available tribal name Haplothripini shown to be appropriate. The character states on which the definition is based are discussed, and a list given of the 34 genera worldwide that can be included. The Australian species in these genera exhibit a diversity of biologies. Three genera involve species that invade galls induced by other thrips: Androthrips monsterae (Moulton) from New Guinea is newly recorded from Australia; Mesothrips jordani Zimmermann from South‐East Asia is newly recorded from Australia, with two new synonyms; the Asia‐Pacific genus, Euoplothrips Hood, includes two species in northern Australia. Three haplothripine genera are associated with grasses: one widespread genus, Apterygothrips Priesner, considered polyphyletic, includes only one Australian species; the only known species of Dyothrips Kudo is Oriental but extends into tropical Australia; Podothrips Hood, a circum‐tropical genus of thrips predatory on grass‐living coccoids, has 11 Australian species, six newly described. One grass‐associated genus, Bamboosiella Ananthakrishnan, is excluded from the Haplothripini. An Oriental genus of leaf‐ and flower‐living species, Dolichothrips Karny, includes one species in northern Australia, D. reuteri (Karny); Membrothrips Bhatti in which this species has been placed is considered a synonym. Karnyothrips Watson includes two species introduced to Australia, both predators of coccoids. Priesneria Bagnall includes three species from Australia, of which one fungus‐feeding species is newly described. Xylaplothrips Priesner is a widespread but ill‐defined, polyphyletic genus that currently includes three little‐known Australian species. Haplothrips is the main focus of this study, and character state variation among the Australian species is discussed. In total 24 species of Haplothrips from Australia are recognised, 14 being new species. Also included are the following, one new generic synonym, four new species synonyms, and one new combination. The 250 species worldwide in the genus are usually associated with the flowers of Asteraceae and Poaceae. In Australia, several of the species are specific to flowers, particularly of Poaceae and Cyperaceae but not Asteraceae. Almost half of these Australian species are presumed to be predatory on other small arthropods, and two have unusual host associations—with salt marsh Chenopodiaceae, and with the sori of Dicksonia tree ferns.     

XLI.—Note on a Megalosaurian tibia from the lower Lias of Wilmcote,Warwickshire

Summary

A new species of Dolichomiris Reuter from India and another from South Africa are described, while costicollis Berg is excluded, bringing the total number of species in the genus to seven. All species are figured and keyed. D. linearis Reuter is recorded from Australia, New Guinea, Indonesia and the New Hebrides. Lectotypes are designated where necessary. A revised key to the genera of Stenodemini is presented.     

LX.—Studies on the Onychophora.—VI. The life-history of Peripatopsis

Summary

Specimens of Notoscolex camdenensis, the type-species of Notoscolex are described for the first time since erection of the genus. The type-species is shown to have stomate megameronephridia median to micromeronephridia in posterior segments, a condition indicating that its affinities lie with a Dichogaster-Megascolides group of genera and not, as has previously been held, with a Megascolex group. Only four other species appear to be congeneric with N. camdenensis. These are endemic in New South Wales, like the type-species, or in Queensland.