Revision of the American Pupisoma species (Gastropoda: Pupilloidea)

The American species of Pupisoma Stoliczka (Gastropoda: Pupilloidea) are revised. Five species are recognized. Pupisoma puella Hylton Scott has been synonymized with P. dioscoricola (C. B. Adams). This species occurs from southern USA through the Caribbean and Central America to the Galapagos Islands and to southern Brazil and northern Argentina. Pupisoma galapagorum Pilsbry, P. bailyi Pilsbry, and P. latens Hylton Scott have been synonymized with P. comicolense H. B. Baker. This species is distributed from Mexico to the Galapagos Islands, southern Brazil, and northern Argentina. Pupisoma costulata sp. n. from Andean forests in Colombia is described as new. The range of P. macneilli (Clapp) extends from southern USA through the Caribbean and Central America to southern Brazil and northern Argentina. A neotype has been designated for P. mediamericanum Pilsbry. Pupisoma michoacanense Pilsbry is a synonym of this species. This species occurs from Mexico to Colombia and on Jamaica.     

Atlantic and Mediterranean species of the genus Andaniexis Stebbing (Amphipoda: Stegocephalidae)

The Atlantic and Mediterranean species of the genus Andaniexis Stebbing, 1906 (Amphipoda: Stegocephalidae) are reviewed, and three new species are described. The species Andaniexis abyssi (Boeck, 1871) which previously was considered a widely distributed species (from Baffin Bay to both Spitzbergen and the Skagerrak), is here divided into three species: Andaniexis abyssi s.str., Andaniexis gracilis n.sp. and Andaniexis lupus n.sp. These species, together with Andaniexis mimonectes Ruffo, 1975, all have allopatric distributions. This is especially evident in northern Norway, where A. abyssi has its northernmost limit: A. abyssi is found in Solbergfjorden (69°10′N), while Andaniexis lupus is found in the adjacent fjord Malangen (69°30′N, the southern limit of this species along the Norwegian coast). Andaniexis eilae n.sp. is described from Iceland. This species is not closely related to the other North Atlantic species of the same genus.     

Observations on the “tenuis group” (Eutardigrada,Macrobiotidae) and description of a new Macrobiotus species

A new species of the tardigrade genus Macrobiotus is described. The species, designated M. ciprianoi n. sp., was isolated from a mixture of Provence broom leaf litter and mosses, and from rock mosses collected in the Sierra de Guadarrama, Madrid (Spain). Given that Macrobiotus ciprianoi n. sp. shares several characters to members of the “tenuis group”, we assessed the taxonomic homogeneity of the group. The new species differs from those of the “tenuis group” according to a unique set of characters related with claw shape, features of the buccal‐pharyngeal apparatus, and egg morphology. Our analysis of holotypes and/or paratypes of “tenuis group” species and other Macrobiotus species with similar characters (M. bondavallii and M. caelicola) reflects the heterogeneity of this group of species as currently described.     

Diversity and distribution of marine Gastrotricha along the northern beaches of the state of São Paulo (Brazil), with description of a new species of Macrodasys (Macrodasyida,Macrodasyidae)

Forty-three putative species were found in this first-ever survey of the marine gastrotrich fauna of Brazil. Samples from 23 locations and 37 sites along the northern coasts of the state of São Paulo yielded 21 species of Macrodasyida (12 genera and five families) and 22 species of Chaetonotida (seven genera and two families). Based on morphology, 26 species appear to be taxa not previously described and thus far endemic to Brazil, 13 are known species, while one in each of the genera Diplodasys, Paradasys and Tetranchyroderma remains undetermined. The overall high diversity contrasts with the generally low number of species from single locations, averaging 4.3 (SD 3.5) spp. per location; very few species were found in sites characterized by very fine sand sediment loaded with detritus and by high water turbidity (e.g. 1 sp. in praia da Fazenda), whereas a well-diversified gastrotrich community was found in localities characterized by medium or fine sand with little detritus and clear water (e.g. 14 spp. in praia de Castelhanos). The macrodasyidans, Paraturbanella sp. 1, Pseudostomella sp. 1 and Tetranchyroderma sp. 1, and the chaetonotidan Heteroxenotrichula sp. 1 were the most common species, often sympatric and particularly frequent and abundant along the continental shoreline (recorded in 50–65% of these localities); on the other hand, Macrodasys fornerise n. sp., Ptychostomella sp. 1, Tetranchyroderma sp. 3 and Thaumastoderma sp. 2, while occurring sometimes in very high numbers, exhibited a much narrower range, being encountered only once or twice during the survey. The main morphometric traits are described of the 13 species considered to be cosmopolitan or with at least an amphi-Atlantic distribution, including Chaetonotus apechochaetus and Halichaetonotus marivagus, reported here for the first time from the Americas; of the 26 species thus far endemic to Brazil, Macrodasys fornerise n. sp., which is characterized by a noticeable 145?µm-long frontal organ, is also described and figured.     

West Indian Neoathyreus and their over-water dispersal (Coleoptera: Scarabaeidae: Geotrupinae)

Neoathyreus androsensis n.sp., is described from Andros Is. in the Bahamas and from Cayo Coco, Cuba. The new species and four other previously described species, now known to occur in the West Indies, are briefly described, keyed and figured. One species, Neoathyreus lanei (Martínez) occurs in the Lesser Antilles and also in northern South America. The four other species occurring on either Cuba, the Bahamas, or Hispaniola are closely related to each other and somewhat less so to the Neoathyreus hamifer complex, known from Mexico and Central America. A minimum of two or three over-water dispersals by ancestral forms seem to be necessary to explain the current distributions of at least some of these West Indian species.     

A synopsis of Martarega White, 1879 (Insecta: Heteroptera: Notonectidae) occurring in the Brazilian Amazonia,with descriptions of three new species

The genus Martarega comprises 15 species and nine of these have been recorded in northern Brazil. Martarega pydanieli sp. nov. from Rondônia, while Martarega nieseri sp. nov. and M. barcelos sp. nov. from Amazonas are described here. Six known species are recorded in the Brazilian Amazonian Region (Amazonas, Pará, Rondônia and Roraima): M. brasiliensis, M. chinai, M. gonostyla, M. membranacea, M. oriximinaensis, and M. uruguayensis. At the outset of this survey the genus Martarega of the Brazilian Amazonian Region held 12 species, but no specimens of M. mcateei, M. hungerfordi, and M. williamsi were collected in the regions sampled. Martarega brasiliensis is newly recorded from Roraima, which comprises the first record of members of this genus in this State. Martarega uruguayensis is newly recorded from Pará and Rondônia, while M. gonostyla from Rondônia. Distinct keys to males and females of Martarega occurring in this region, including these new species, are provided.     

The pygmaeus-group of Euops Schoenherr (Coleoptera,Curculionoidea, Attelabidae), weevils associated with Nothofagus in New Guinea

A new species group of Euops Schoenherr from the highlands of New Guinea is described and referred to as the ‘pygmaeus-group’. It is comprised of 14 species which, except for E. singularis VOSS, are described as new: E. anggiensis sp. n., E. bicolor sp. n., E. convexus sp. n., E. fraterculus sp. n., E. ibelensis sp. n., E. kurulu sp. n., E. nothofagi sp. n., E. parvus sp. n., E. paraconvexus sp. n., E. platyrostris sp. n., E. porulosus sp. n., E. pygmaeus sp. n., and E. sedlaceki sp. n.. Most of the species are of relatively small body size. They form a monophyletic group characterized by the synapomorphic loss of the ventral suture between the galea and palpifer of the maxilla. Two other groups are recognized, the eucalypti-group and the quadrifasciculatus-group. They are believed to be closely related to the pygmaeus-group and all three species groups together are referred to as the ‘pygmaeus-complex’. A cladistic analysis of the pygmaeus-group is performed. All species with host records are reported to feed on the fresh leaf flush of Nothofagus. This is the first record of a group of leaf-chewing arthropods restricted to the subgenus Brassospora of Nothofagus.     

New species of Orasema (Hymenoptera: Eucharitidae) from Central and South America

ABSTRACT

Six new species of Orasema are described. Four species are placed into a newly recognised lasallei species group: Orasema lasallei sp. nov. (Costa Rica, Honduras, Mexico, Nicaragua), O. janzeni sp. nov. (Costa Rica), O. peckorum sp. nov. (Brazil) and O. vasquezi sp. nov. (Colombia). One species is placed in the wayqecha group, Orasema masneri sp. nov. (Ecuador). One other species is currently unplaced to species group, O. longinoi sp. nov. (Costa Rica). Orasema lasallei was taken from a nest of Pheidole vorax (Fabr.) and an undescribed species near O. longinoi from a nest of Pheidole dossena Wilson (Myrmicinae). The morphology of the digitate labrum is discussed with respect to other Eucharitidae.     

Notes on the Amazonian genus Hyogonia China (Hemiptera: Cicadellidae: Proconiini) with a description of a new species

Oncometopia batesi Distant, described from northern Brazil (State unknown), is transferred to the genus Hyogonia China. Hyogonia brasiliana sp. nov. is described from Northern Brazil, Pará State, the easternmost record of the genus. The new species is easily distinguished from all other Hyogonia species by the following combination of characters: (1) pronotum without distinct transverse contrasting band; (2) pygofer with short processes that do not attain its apex; (3) both pairs of aedeagal atrial processes divergent and short, never longer than aedeagal shaft; and (4) aedeagal caudal pair of atrial processes with bases close to each other, appearing as a single bifurcated robust process. Hyogonia batesi and H. brasiliana, together with an unidentified female from Acre State, represent the first authentic records of the genus Hyogonia from Brazil. Previous records of H. reticulata (Melichar) and H. youngi Emmrich and Lauterer from Brazil are erroneous as their recorded locality ‘Cumbase’ is most probably located in Peru (San Martín Department). Other new records from countries and major federal divisions are noted as follows: the genus Hyogonia (unidentified species) from Barinas State (Venezuela); H. batesi from Ecuador (Orellana Province); H. reticulata from Amazonas and Junín Departments (Peru); and H. youngi from Ecuador (Napo Province). A key to the known species of Hyogonia is provided.     

Psocoptera (Insecta) from Bermuda

Seventeen species of Psocoptera are recorded from Bermuda, increasing the known number from three. Four new species are described: Echmepteryx atlantica n. sp., Caecilius bermudensis n. sp., Indiopsocus hilburni n. sp., and I. nebulosus n. sp. The latter two species appear to be closely related and two alternative hypotheses are proposed for their origin. Ectopsocus spilotus Thornton and Wong is recorded outside the central Pacific area for the first time. The species are assigned to four categories according to probable duration of time on Bermuda and probable source of original invaders: (1) endemics (four species); (2) species with wide distributions in the tropics and subtropics, not restricted to islands and sea coasts (four species); (3) species largely restricted to sea coasts and islands, not necessarily tropical (three species); (4) species widely distributed in southeastern United States, not restricted to the sea coast or the subtropical region (five species).     

The genus Monacha in the Western Caucasus (Gastropoda: Hygromiidae)

Seven Monacha species have been found in the Western Caucasus region. Six of these species are endemic there. They all belong to the subgenera Paratheba and Metatheba. Three species are new to science. M. (Paratheba) claussi, sp. n. and M. (Metatheba) subcarthusiana (Lindholm) are the only Monacha species, in which the right ommatophoral retractor sometimes crosses with the penis. This might be due to an ancestral polymorphism, atavism or a basal position of the two species within Monacha, which would mean that Paratheba is paraphyletic and Metatheba is polyphyletic. Monacha (Monacha) cartusiana (O. F. Muller), the only Monacha sensu stricto species in the Caucasus region, has probably been introduced by man. The shell and the genitalia of every species are described and illustrated. The synonyms and all Caucasian locality records are listed. The distributions of the species are illustrated on UTM-grid maps.     

Hysterothylacium,Iheringascaris and Maricostula new genus,nematodes (Ascaridoidea) from Australian pelagic marine fishes

The state of knowledge of Australian fish parasitic ascardioids is briefly reviewed. The following species are described in full: Hysterothylacium cornutum, H. pelagicum, H. scomberoidei n. sp. Additional data are provided for Iheringascaris inquies. Maricostula n. gen. is established for those species that have lips with lateral flanges, deep postlabial grooves, interlabia, and elongate intestinal caecum, pyriform ventriculus, long sac-like ventricular appendix, an expanded filamentar excretory system and excretory pore near the nerve ring. The males of all the species have precloacal crests. Species included are: M. caballeroi (Deloya), M. cenatica n. sp., M. histiophori (Yamaguti), M. incurva (Rudolphi), M. makairi n. sp. and M. tetrapteri n. sp. All are recorded from billfishes, and all but M. caballeroi are recorded from Australian waters. The systematic position of the genus is discussed. New characters of taxonomic utility used in the discrimination of fish parasitic ascaridoids are discussed. These include fine details of alae, male caudal crests and papillae. We demonstrate that these characters can be used to separate species of Maricostula. Remarks are also given on nomenclatural problems within the Indo-Pacific fish parasitic Ascardioidea.     

Chelodesmid studies. XVIII. On some new or poorly-known taxa in the tribe Batodesmini (Polydesmida: Chelodesmidae)

Summary

Some new and poorly-known taxa of batodesmine diplopods are described from northern South America, with notes on their affinities. New genera include Igaraparana, based on I. batesi, sp. nov. from Amazonian Colombia; Curimagua, based on C. granulata, sp. nov. from Edo. Falcon, Venezuela; and Cheirogonus, based on C. pittieri, sp. nov. from Panama. The inadequately proposed species Leptodesmus interrupticolor Silvestri (southeastern Ecuador) is redescribed and figured from type material, and referred to Cordilleronomus as its third species. The enigmatic genus Tunochilus Chamberlin, 1950, is clarified and referred to the Batodesmini from its incorrect location in the Cryptodesmidae, with notes and drawings made from a paratype of the type species T. marginis; the presumptive congener T. marcuzzii is described as new from material taken at Caracas, Venezuela.     

New data on Hydrobiidae systematics: two new genera from the Iberian Peninsula

The taxon Caprella acanthifera auct., usually considered to be one ‘highly variable’ species, is shown here to consist of a number of closely related species. In this paper three species of the group lacking an axillary spine near the insertion of gnathopod 2 are described and illustrated; the taxa with an axillary spine will be dealt with in a separate paper. This paper redefines Caprella acanthifera Leach 1814, sensu stricto; the nominal taxa C. hystrix, C. calva, C. aspera, and C. leptonyx are considered to be junior synonyms. Also described are C. stella sp.n., from starfish at the Azores and C. cavediniae sp.n., living among algae in the Mediterranean. The sources of apparent morphological variability in caprellids are discussed.     

Description of new Ceratitis MacLeay (Diptera,Tephritidae) species from Africa

As an addition to earlier published systematic revisions of the genus Ceratitis MacLeay, five new species are hereby described: Ceratitis (Pardalaspis) millicentae sp. n., C. (Ceratalaspis) oraria sp. n., C. (C.) perisae sp. n., and C. (C.) perseus, sp. n. all from Kenya, and C. (C.) ealensis sp. n. from the Democratic Republic of Congo. Their relationship with closely allied species within the respective subgenera is discussed and differentiating characters given.     

A new genus of Neotropical whitefly,secreting blue-iridescent wax (Sternorrhyncha,Aleyrodidae, Aleurodicinae), and its parasitoids (Hymenoptera,Aphelinidae)

Azuraleurodicus pentarthrus Martin gen. and sp. n. is described from three different host plant families in Central and northern South America. Adults of both sexes were reared in culture and adult characters are discussed in addition to those of puparia. The systematic position of Azuraleurodicus within the Aleurodicinae is discussed. One new combination, Aleuronudus ferrisi (Sampson and Drews) comb. n., is here proposed in the course of discussion. The secretion of blue wax by whiteflies is discussed and illustrated by colour photographs. Two species of hymenopterous parasitoids, Dirphys aphania Polaszek sp. n. and Encarsiella pithecura, Polaszek sp. n. emerged from puparia of Azuraleurodicus pentarthrus and are here described.     

A troublesome sibling species complex of scuttle flies (Diptera: Phoridae) revisited

The complex of species resembling Megaselia pulicaria (Fallén) has proved to be one of the most troublesome challenges to attempts to resolve the recognition of species within the huge genus Megaselia Rondani. The new data presented allow more confident recognition of species and reveal that most published records represent misidentifications. The penis complex is discussed and its use in species identification demonstrated. All specimens currently assigned to M. pulicaria, other than the lectotype, are consequently reassigned to other species. These include the reinstatement of M. angusta (Wood) and its newly designated subspecies M. angusta angustina Schmitz, M. eupygis Schmitz, M. longifurca (Lundbeck) and M. monochaeta (Strobl) as valid species, and the recognition of M. buchsi sp. nov. from Germany, M. iberiensis sp. nov. from Spain, M. martensi sp. nov. from Nepal, M. nectergata sp. nov. from Britain, M. oviaraneae sp. nov. from Britain, and M. rotundapicis sp. nov. from Scotland. Five synonyms are proposed. The hitherto unknown male of M. petraea Schmitz is described. It and M. longifurca are reported from Britain and the latter from the Nearctic Region also. Natural history information is summarized for each species. In particular, it is reported that the species whose larvae prey upon spider eggs are M. angusta, M. longifurca and M. oviaraneae.     

A revision of Tachyphron Brown and description of two new genera within the Ariphron group (Hymenoptera: Tiphiidae)

The Australasian genus Tachyphron Brown is revised, and two new New Guinean genera, Deuterothynnus gen. n. and Heligmothynnus gen. n., described. Fifteen species are included in Tachyphron of which seven are described as new (T. aculeatus sp. n., T. athertonensis sp. n., T. mantonensis sp. n., T. neosubfragilis sp. n., T. nigrisetatus sp. n., T. planus sp. n. and T. townsvillensis sp. n.), and a further two, Aelurus comatus Smith and Aelurus fragilis Smith, transferred from Tachynomyia. Thynnus insularis Smith is transferred from Tachynomyia Guérin to Deuterothynnus and a further three new species, D. fulvicentratus sp. n., D. fulvisetatus sp. n. and D. parallelus sp. n. described. Aelurus atratus Cameron and Takyomyia sabronensis Kimsey are transferred from Tachynomyia and Tachyphron, respectively, to Heligmothynnus and a further two new species, H. microspinus sp. n. and H. neoaratus sp. n. described. T. megacephala (Turner) is synonymized with Tachynoides flavopicta (Ritsema) and, therefore, removed from Tachyphron. A key to the Ariphron group of genera is provided, as well as keys to the males of Tachyphron, Deuterothynnus and Heligmothynnus, although the male of D. insularis is associated tentatively with the female holotype only on the basis of collection records. Only the females of D. insularis, T. armidalensis Brown and T. subtriangularus Brown are known, the latter two being described for the first time. Evidence is presented to suggest that T. subtriangularus may be bivoltine in northern Australia.