Palaearctic species of Ammoplanus (Hymenoptera: Sphecidae)

The Palaearctic species of the genus Ammoplanus Giraud are revised, based on material of over 1750 specimens from Europe, North Africa and Central Asia. For almost all species it was possible to examine the type material and in six species the lectotypes have been designated, in two cases (A. perrisi and A. marathroicus) the neotypes. As a result 38 species are recognized as valid, keyed out and commented on. Eight species are described as new and four species names are here placed in synonymy. Several species are of wide distribution, which made it necessary to include also those described from Central Asia and Mongolia. The keys are made to all Palaearctic species but no material is known from the regions east of Mongolia, although four undescribed species are available from South-East Asia. The Afrotropical species are listed and most of them were also examined. Two subgenera are recognized in the region; 32 species are classified in Ammoplanus sensu stricto and six in the subgenus Ammoplanellus. In the nominal subgenus the following species are described as new: A. atlasensis from Morocco, A. biscopula from Israel, A. biskrensis from Algeria, A. denesi from Turkey (Anatolia), A. freidbergi and A. kaplanae from Israel, A. minutus from Bulgaria and Turkey and A. strumae from Bulgaria and Greece. New synonymy: A. angelae becomes a junior synonym of rhodesianus (known before only from Africa), A. crudelis a synonym of A. kaszabi, A. handlirschi a synonym of A. marathroicus, A. suarezi a synonym of A. kohlii, A. subcompressus of A. gegen, A. orbiculatus of A. simplex and A. zarcoi of A. kohlii; A. wesmaeli is confirmed as a synonym of A. perrisi. Otherwise the synonymy is corrected under A. bischoffi and the names A. dusmeti and A. curvidens are re-established as valid. In a short review of the African species A. (Ammoplanellus) cradockensis Arnold, 1959 is placed as a new synonym under A. (Ammoplanellus) latiscapus Leclercq, 1959. The nomenclatural changes are explained under the relevant species.     

Habitat selection and nest construction behaviour in some Afrotropical species of Ammophila (Hymenoptera: Sphecidae)

Nest construction behaviour and nest site selection are described for seven species of Afrotropical Ammophila.

Four species were atypical in their nest siting. Ammophila beniniensis selected shaded sites in well vegetated habitats, A. vulcania selected small clumps of vegetation in otherwise open habitats. A. insignis nested in vertical banks, old animal burrows and caves. A. braunsi is unique amongst Ammophila in its use of abandoned burrows of other wasps in non-friable clay soils. A. ferrugineipes, A. dolichodera and A. dolichocephala nested mostly in open habitats.

Methods of soil waste disposal, sealing of nests and their final coverage, differed interspecifically, in some cases intraspecifically, but often apparently in response to the habitat.

Temperature conditions, affected by habitat, influenced the depth to which nests were dug by some species. A. insignis switched sites in response to seasonal changes in temperature regimes in vertical banks.

The existence in particular species of both primitive and advanced aspects of nesting behaviour questions their use in assessing the evolutionary status of these species.     

Nesting behaviour and prey of Pseudoplisus Ashmead (Hymenoptera: Sphecidae)

Nesting behaviour and prey of two species of Pseudoplisus, a gorytine wasp genus in the sphecid subfamily Nyssoninae, are described. P. natalensis is a solitary nester in southern Africa nesting often in the soil in flower-pots. P. ranosahae in Madagascar nests in the ground in large aggregations. Both species show a high degree of prey specificity. Nests are provisioned only with large adult froghoppers (Aphrophoridae), P. natalensis preying on Ptyelus grossus and P. ranosahae on Ptyelus goudoti.     

Two new species of Cephalastor Giordani Soika (Hymenoptera: Vespidae: Eumeninae), with a review of the phylogeny of the genus

Cephalastor Giordani Soika is a small Neotropical genus of eumenine wasps comprising 12 species, most of them described at the beginning of this century. The genus was revised by Garcete-Barrett, and a cladistic analysis carried out. Cephalastor humeralis n. sp. and C. minarum n. sp. are described herein, and the phylogenetic relationships among the species of the genus inferred from morphological data. A total of 34 characters were constructed (an increase of more than 209% from Garcete-Barrett's 2003 Garcete-Barrett, BR. 2003. La taxonomía del género Cephalastor Soika (Hymenoptera: Vespidae: Eumeninae), parte II. Bol Mus Nac Hist Nat Parag., 14(1–2): 80–97.  [Google Scholar] study) and new taxa added as outgroups. The monophyly of the genus was corroborated, and the relationships were stable under both equal and implied weighting schemes. New collecting records are added for Cephalastor bossanova Garcete-Barrett and Cephalastor estela Garcete-Barrett.     

Notes on the natural history of Stizus perrisii ibericus Beaumont (Hymenoptera: Sphecidae)

A study of the biology of Stizus perrisii ibericus revealed some aspects about the behaviour of males, the nesting behaviour of females, and larval development. The slow mass provisioning of this species is compared with the provisioning behaviour of S. continuus and S. pulcherrimus. Cocoons are constructed in seven stages although, compared with other species, there are differences in the duration of each stage. Natural enemies of the wasp include miltogrammine flies, several cuckoo wasps, and velvet ants.     

Taxonomic review of Coccobius species (Hymenoptera: Aphelinidae) from China,with notes on their use in biological control of scale insect pests (Hemiptera: Diaspididae), and description of a new species

The Chinese species of the genus Coccobius Ratzeburg (Hymenoptera: Aphelinidae) are reviewed. Sixteen species, C. abdominis Huang, C. annulicornis Ratzeburg, C. azumai Tachikawa, C. chaoi Huang, C. curtifuniculatus Huang, C. flaviceps (Girault & Dodd), C. flavicornis Compere & Annecke, C. fulvus (Compere & Annecke), C. furviflagellatus Huang, C. furvus Huang, C. languidus Huang, C. longialatus Huang, C. longifuniculatus Huang, C. maculatus Huang, C. wuyiensis Huang, and one new species, Coccobius albiscutellum Wang and Huang sp.nov., are treated. A key is provided for the recognition of the females of all Chinese species of Coccobius, and notes are given on their use in biological control of scale insect pests. http://www.zoobank.org/urn:lsid:zoobank.org:pub:F510059B-F176-46BA-B7B5-8252B06BE91F     

The Old World genera of braconine wasps (Hymenoptera: Braconidae)

An illustrated key is provided to the Old World genera of Braconinae. In addition, notes are provided on each genus and on a number of characters employed in the key. One new tribe and five new genera based on two existing and three new species are described. The new taxa are as follows: Angustibracon gen. nov. from India (type-species: Bracon leptogaster Cameron, 1899), Bathyaulacini tribus nov. (type-genus: Bathyaulax Szépligeti, 1906), Cassidibracon gen. nov. from West Africa (type-species: Cassidibracon castus sp. nov.), Serraulax gen. nov. from the Afrotropical Region (type-species: Iphiaulax lukombensis Cameron, 1912), Virgulibracon gen. nov. from Australia (type-species: Virgulibracon vulsus sp. nov.) and Vomeribracon gen. nov. from New Guinea (type-species: Vomeribracon amplipenniger sp. nov.). The Pseudodicrogeniini Achterberg, 1976 is synonymized with the Euurobraconini Ashmead, 1900. Ten new generic synonymies are reported: Atanycolimorpha Viereck, 1913 (= Cyanopterus Haliday, 1835), Caenoprymnus Cameron, 1909 (= Soter Saussure, (1890) 1892), Coelodontus Roman, 1912 (= Trigastrotheca Cameron, 1906), Cynodobracon Cameron, 1912 (= Nedinoschiza Cameron, 1911), Iphioilta Ramakrishna Ayyar, 1928 (= Chaoilta Cameron, 1899), Kenema van Achterberg, 1983 (= Trigastrotheca Cameron, 1906), Odontopygia Enderlein, (1918) 1920 (= Trigastrotheca Cameron, 1906), Paraspinaria Cameron, 1905 (= Mesobracon Szépligeti, 1902), Pseudoglyptomorpha Tobias, 1957 (= Pseudovipio Szépligeti, 1896) and Glyptomorpha (Remorpha) Shestakov, 1932 (Pseudovipio Szépligeti, 1896).     

Two new species of Clinocentrus Haliday (Hymenoptera: Braconidae) from Argentina,with notes on South American species

The Argentine species of the genus Clinocentrus Haliday are revised. Two new species are described, Clinocentrus mamull sp. nov. and Clinocentrus roigi sp. nov., which represent the southernmost records of the genus in South America. Additionally Clinocentrus nigripectus Enderlein is redescribed and a key to known South American species is provided.     

Revision of the Neotropical species of the tribe Euderomphalini (Hymenoptera: Eulophidae)

The Neotropical species and genera of the tribe Euderomphalini (Eulophidae: Entedoninae) are revised. Five genera are described as new: Cabeza, including six species (baeostigma (§, ¦), canaliculata (§), laticeps (¦), petiolata (§), planiscapus (§), ugaldei (§)); Itahipeus, including two species (brasilicola (§, ¦), euryceps (¦)); Monterrondo, including one species (aphelosoma (§)); Sifraneurus, including one species (strigifer (§)); Xenopomphale, including one species (sulcata (§)); all above-mentioned species are described as new. Five additional genera are included: Aleuroctonus LaSalle and Schauff, with four species, one described previously: vittatus (Dozier) (§, ¦), and three described here: latiscapus (§), marki, (§, ¦) and metallicus (§); Entedononecremnus Girault, with 14 species, two described previously: krauteri Zolnerowich and Rose (§, ¦), unicus Girault (§, ¦), and 12 described here: bennetti (§, ¦), bimaculatus (§, ¦), convexus (§, ¦), crassicornis (§, ¦), depressus (§), fulgens (§), hansoni (§, ¦), imdasus (§, ¦), parfer (§, ¦), reticulatus (§), tripar (§) and unicarinatus (§); Euderomphale Girault, with two species, one described previously: flavimedia (Howard), and one described here: sulciscapus (§, ¦); Neopomphale LaSalle and Schauff, with 20 species, two described previously: aleurothrixi (Dozier) (§, ¦), quercicola (Dozier) (§, ¦), and 18 described here: azofeifai (¦), cerrobius (¦), depilis (§), dichrous (§, ¦), erecta (¦), graciliclava (§), longicornis (¦), longipilis (§, ¦), nonaequa (§, ¦), noyesi (§), pinguicornis (§, ¦), prymna (§), rubescens (§, ¦), rubii (§), silvestris (¦), transversa (§), umbonata (§, ¦), xenipennis (§); Dasyomphale LaSalle and Schauff, including one species (chilensis LaSalle and Schauff). The genera are placed in either one of three genus-groups, one of which is newly created. The species Euderomphale quercicola Dozier is newly combined to Neopomphale. Lectotypes are designated for Entedononecremnus unicus Girault, Euderomphale quercicola Dozier and Gyrolasia flavimedia Howard. Key to genera and species are included.     

A new neobisiid pseudoscorpion species from Crete (Greece), with notes on its morphology,distribution, evolution,and phylogeny

This study, based on an analysis of some palaeo‐Mediterranean pseudoscorpions (Ernstmayria venizelosi n. sp. and E. apostolostrichasi ?ur?i? and Dimitrijevi? from the family Neobisiidae) represents an outstanding improvement of our knowledge of the diversity of relict and distributionally limited false scorpions from the ancient (or proto‐) Aegean area. Both E. venizelosi n. sp. and E. apostolostrichasi originated in the proto‐Balkan region; their ancestors are to be sought among extinct forms. Since Ernstmayria species presently inhabit the ‘peripheral’ location of the ancient Aegeis, it is evident that their remote ancestor gave rise to a number of recent genera which presently constitute two generic groups inhabiting the Mediterranean region: Neobisium Chamberlin, Occitanobisium Heurtault, Roncobisium Vachon, Protoneobisium ?ur?i?, and Pennobisium ?ur?i?, on the one hand, and Acanthocreagris Mahnert, Balkanoroncus ?ur?i?, Insulocreagris ?ur?i?, and Roncocreagris Mahnert, on the other.     

Six new species of Platygastrinae from the British Isles (Hymenoptera,Platygastridae)

Six species of Platygastrinae new to science are described and illustrated, namely Platygaster anglica, P. nixoni, P. sublongicornis, Synopeas londiniense, S. nottoni, and S. noyesi.     

A review of the genus Geraldus Fitton (Hymenoptera: Ichneumonidae: Banchinae), with description of a new species

An undescribed species of Banchinae from Chile in the Natural History Museum collections is described as Geraldus fittoni Broad, sp. nov. Four more specimens of Geraldus cambrensis have been found, from Chile and Argentina. The monophyly of Geraldus and of Philogalleria are supported.     

Two new species of Hymenoepimecis (Hymenoptera: Ichneumonidae: Pimplinae) with notes on their spider hosts and behaviour manipulation

The wasp genera included in the Polysphincta genus-group develop exclusively as koinobiont ectoparasitoids of spiders, an unusual pattern within Hymenoptera. All known species of Hymenoepimecis attack orb-web spiders and Hymenoepimecis argyraphaga was previously described inducing a sudden modification of its host's web-building behaviour. Information on parasitoid–host interactions and even on host identities for most members of the genus-group, however, remain scarce. In this study we describe two new species of Hymenoepimecis and present information on their hosts, spiders of the genera Leucauge (Tetragnathidae) and Manogea (Araneidae). Cocoon webs of both host species are distinct from their normal webs. The modified cocoon web of Leucauge roseosignata constructed under the influence of Hymenoepimecis japi sp. nov. is very similar to the cocoon web described for Leucauge argyra. It is composed of three axes, including several radial threads each, and a hub used by the larva to attach the cocoon's suspension line. Spiral viscous threads, present in normal webs and in webs of parasitized spiders carrying larvae in the initial instars, are absent in cocoon webs. In the cocoon web of Manogea porracea the horizontal sheet is poorly structured and the cocoon is attached far from the spider's normal resting position, in an area composed of a densely tangled structure.     

Taxonomy and cladistic analysis of the subgenus Xenomorellia Malloch (Diptera: Muscidae: Morellia Robineau-Desvoidy) with description of two new species

Xenomorellia Malloch, a subgenus of Morellia Robineau-Desvoidy, is revised to include two new species, Morellia (Xenomorellia) inca Nihei and Carvalho sp. nov. from South America, and M. (X.) maia Carvalho and Nihei sp. nov. from Costa Rica and Mexico. Diagnoses for M. (X.) holti (Malloch) and M. (X.) montanhesa (Albuquerque) are provided, as well as an identification key to the four species of the subgenus. A cladistic analysis was performed to test the monophyly of Xenomorellia and to recover the phylogenetic relationships among its species. Tree searches resulted in one single most-parsimonious cladogram, wherein the monophyly of Xenomorellia is supported, as well as a sister-group relationship with the Neotropical subgenus Trichomorellia Stein. Xenomorellia was divided into two clades: one with Caribbean–Andean species (maia?+?inca), and another with species from southeastern South America (holti?+?montanhesa).     

The African species of Zaglyptogastra (Hymenoptera) with a checklist of world species

An illustrated key is provided to the African species of the braconine wasp genus (Zaglyptogastra Ashmead. Four new species are described: Z. afenestrata sp. nov. from Ruanda, Z. levisulcata sp. nov. from Uganda, Z. tenuicauda sp. nov. from Angola, Congo and Zaire, and Z. virgulivena sp. nov. from Zaire. A replacement name, Z. caudatula nom. nov. is proposed for Calliidia caudata Szépligeti, 1914b. Six new synonymies are proposed: Zaglyptogastra basalis (Szépligeti; 1914) (= Z. equitator (Wiedemann, 1824)); Z. leucogaster (Cameron, 1909a) (= Z. nigripennis (Szépligeti, 1908)); Z. stanleyi (Cameron, 1912), Z. maxima (Szépligeti, 1914b), Z. eleganta (Shenefelt, 1978) and Z. tomentosa (Fahringer, 1931) (= Z. tincticanaliculata (Cameron, 1912)). Notes are provided on the African species and species groups. A checklist of world species is given.     

Cornutorogas,a new genus with four new species of the subfamily Rogadinae (Hymenoptera: Braconidae) from the Oriental region

A new genus, Cornutorogas gen. nov., and four new species, Cornutorogas orientalis Chen and Belokobyskij, sp. nov. (China, Vietnam), C. maetoi van Achterberg, sp. nov. (Thailand), C. javensis van Achterberg, sp. nov. (Java) and C. sumatrensis van Achterberg, sp. nov. (Sumatra), of the subfamily Rogadinae (Hymenoptera: Braconidae) from the Oriental Region are described and illustrated. The placement of this genus in the Rogadini is discussed.     

Notes on some palaearctic and oriental Systropha,with descriptions of new species and a key to the species (Hymenoptera: Apoidea: Halictidae)

An annotated list of the palaearctic and oriental species of the genus Systropha Illiger, 1806, includes additional records for some less well-known species. A lectotype is designated for Systropha hirsuta Spinola, 1839. Systropha christae Warncke, 1992, is synonymized with S. hirsuta. Systropha androsthenes and S. diacantha, spp. n., are described from the Gulf States. A key for the separation of the males is given.