Differing morphology and ultrastructure of the male copulatory apparatus in species-groups of Anagrus Haliday (Hymenoptera: Mymaridae)

Male genitalia of 13 members of the mymarid wasp genus Anagrus have been studied: Anagrus (Anagrella) mymaricornis (Bakkendorf), Anagrus (Paranagrus) unilinearis Soyka, A. (Anagrus) ustulatus Haliday, A. (A.) erythroneurae Trjapitzin and Chiappini, A. (A.) atomus (Linnaeus), A. (A.) aegyptiacus Soyka, A. (A.) vilis Donev, A. (A.) sensillatus Viggiani and Jesu, A. (A.) breviphragma Soyka, A. (A.) incarnatus Haliday, A. (A.) obscurus Förster sensu Soyka, A. (A.) fennicus Soyka and A. (A.) avalae Soyka. Taxonomically significant differences between subgenera and the atomus and incarnatus species-groups are described. The functional significance of various features, and in particular, the arrangment of sensilla are discussed.     

A review of the encyrtid wasp tribe Aenasiini,with descriptions of new Afrotropical taxa (Hymenoptera: Chalcidoidea)

The tetracnemine tribe Aenasiini is discussed and a key is provided for the separation of the nine genera that can be attributed to this tribe. Neodiscodes is synonymized with Aenasius and the following seven species, which were originally assigned to the former genus, are transferred to Aenasius as new combinations: abengouroui (Risbec), comperei (Kerrich), lepelleyi (Kerrich), martinii (Compere), indicus (Narayanan & Subba Rao), parvus (Kerrich) and subbaraoi (Kerrich). A key is given for the separation of the four Afrotropical species of Aenasius and the male of A. comperei is described for the first time. Incisencyrtus gen. n. is described for the following three new Afrotropical species: secus (the type-species), sirus and afer. Cladiscodes incisius sp. n. is described from South Africa and it represents the first species of this genus from Africa. New distribution records are given for three Alfrotropical species of Metaphaenodiscus.     

Two new species and additional distributional records of Acmopolynema Ogloblin (Hymenoptera: Mymaridae) from India

Two new species of Acmopolynema Ogloblin (Hymenoptera: Mymaridae), Acmopolynema pteron Manickavasagam & Palanivel sp. nov. and Acmopolynema pseudotachikawai Manickavasagam & Palanivel sp. nov., are described from India; a revised key to Indian species of the genus is provided. Acmopolynema shrawastianum Hayat & Anis syn. nov. is synonymized under Acmopolynema indochinense (Soyka); Acmopolynema orchidea Triapitsyn & Berezovskiy is reported from India, and new distributional records of Acmopolynema tachikawai Taguchi from Brunei and India are also given.www.zoobank.org/urn:lsid:zoobank.org:pub:3098457A-CA39-49CD-AEF1-19E788C33B78     

A taxonomic review of the genus Exochus Gravenhorst (Hymenoptera: Ichneumonidae: Metopiinae) from South Korea with descriptions of ten new species

The South Korean species of the genus Exochus are revised. Ten new species, Exochus acostulatus Lee & Choi, sp. nov., Exochus adentatus Lee & Choi, sp. nov., Exochus areolaris Lee & Choi, sp. nov., Exochus carinalis Lee & Choi, sp. nov., Exochus dentisternum Lee & Choi, sp. nov., Exochus depressus Lee & Choi, sp. nov., Exochus nigritulus Lee & Choi sp. nov., Exochus occipitalis Lee & Choi, sp. nov., Exochus orbitalis Lee & Choi, sp. nov. and Exochus propodealis Lee & Choi, sp. nov., are described. Also, 21 species of this genus are reviewed and newly recorded from South Korea, with diagnoses provided. A key to the South Korean species of Exochus and illustrations of external characters are provided.

http://zoobank.org/urn:lsid:zoobank.org:pub:B28700A7-9CA7-4AE7-9816-9C8F4CD46160     

First record of the genus Brachyzapus Gauld and Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam,with descriptions of six new species

Based on a recent collection of ichneumonids from Lao Cai Province, at an elevation above 1500 m above sea level, the genus Brachyzapus Gauld and Dubois, 2006 Gauld, ID and Dubois, J. 2006. Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae: a taxonomic revision of spider ectoparasitoids. Syst Entomol., 31: 529–564. [Crossref], [Web of Science ®] , [Google Scholar] is reported from Vietnam for the first time and six new species are described: B. carinatus sp. nov., B. convexus sp. nov., B. duboisi sp. nov., B. fansipanensis sp. nov., B. hoanglienensis sp. nov. and B. politus sp. nov. Notes on distinguishing characters are provided and the new species are included in a tentative key to 12 species of Brachyzapus. Three species are formally transferred from Zabrachypus to Brachyzapus: B. atripedalis (Sheng, 2001), B. nitidus (Hao and Sheng, 2002 Hao, D and Sheng, ML. 2002. A new species of genus Zabrachypus Cushman (Hymenoptera: Ichneumonidae). J Northeast Forestry Univ., 30: 95–96.  [Google Scholar]) and B. nonareaeidos (Wang, 1997) are new combinations.     

Review of Loxopus Townes (Hymenoptera,Ichneumonidae, Cryptinae), with descriptions of six new species

The taxonomic limits of Loxopus Townes are reviewed. The genus is characterized by the lateral margin of the clypeus projecting as subtriangular lobe; lateral lobes of fourth tarsomeres distinctly longer than mesal lobes; fore wing vein 3r-m absent; hind wing vein 2-1A absent or vestigial; and first metasomal tergite with a basolateral tooth. It occurs from Mexico to southern Brazil. Nine species are recognized, of which six are described as new: L. dodecius Santos et Aguiar, L. duckei Santos et Aguiar, L. exius Santos et Aguiar, L. ichilus Santos et Aguiar, L. tenuis Santos et Aguiar, and L. venezuelanus Santos et Aguiar. The type species, L. australis Townes, and L. multicolor Kasparyan et Ruíz-Cancino are redescribed, and a diagnosis is provided for L. unicolor Kasparyan et Ruíz-Cancino, not examined. All studied species are illustrated and distribution records are mapped. An identification key for the species of Loxopus is presented.

http://zoobank.org/urn:lsid:zoobank.org:pub:5B73E8B4-1288-4FD9-AABD-845B5085FFE3     

A review of the genus Geraldus Fitton (Hymenoptera: Ichneumonidae: Banchinae), with description of a new species

An undescribed species of Banchinae from Chile in the Natural History Museum collections is described as Geraldus fittoni Broad, sp. nov. Four more specimens of Geraldus cambrensis have been found, from Chile and Argentina. The monophyly of Geraldus and of Philogalleria are supported.     

Revision of Aleiodes (Hemigyroneuron) parasitic wasps (Hymenoptera: Braconidae: Rogadinae) with reappraisal of subgeneric limits,descriptions of new species and phylogenetic analysis

The parasitic rogadine braconid wasp subgenus Hemigyroneuron, previously and inappropriately synonymized with Aleiodes (Chelonorhogas) as a result of misidentification, is here synonymized with Aleiodes (Pholichora). A key to the 26 species of Aleiodes (Hemigyroneuron) is presented, including 16 which are described and illustrated as new: A. (H.) amoretae Butcher and Quicke, A. (H.) faddenae Butcher and Quicke, A. (H.) glandularis Butcher and Quicke and A. (H.) plurivena Butcher and Quicke from South Africa; A. (H.) bakeri Butcher and Quicke from Indonesia; A. (H.) bengalensis Butcher and Quicke from India; A. (H.) cusackae Butcher and Quicke from Uganda; A. (H.) elgon Butcher and Quicke, A. (H.) meruensis Butcher and Quicke and A. (H.) pappi Butcher and Quicke from Kenya; A. (H.) fenwickae Butcher and Quicke from Congo; A. (H.) pseudospeciosus Butcher and Quicke from Brunei; A. (H.) roberti Butcher and Quicke from Thailand; A. (H.) sharkeyi Butcher and Quicke from Madagascar; A. (H.) trianguliscleroma Butcher and Quicke from Malawi and Kenya; and A. (H.) ugandaensis Butcher and Quicke from Uganda. A. (H.) apicale (Brues) comb. nov., A. (H.) dubiosus (Fullaway) comb. nov., A. (H.) speciosus (Baker) comb. nov. and A. (H.) suffusus (Baker) comb. nov. are redescribed and illustrated. Males of most species of A. (Hemigyroneuron) are shown to possess a glandular pore opening near the mid-posterior margins of metasomal tergites 4–6, though the situation in A. (H.) dubiosus is different. Results of morphological phylogenetic analysis are presented and indicate clear separation between African and Asian species groups. Two of the new species have been reared from Geometridae and one apparently from Pieridae.     

The species of Astymachus Howard (Hymenoptera: Encyrtidae), potentially important parasitoids of Aclerdidae (Hemiptera: Coccoidea) associated with grasses (Poaceae), with descriptions of three new species

ABSTRACT

Species of Astymachus (Hymenoptera: Encyrtidae) are of potential economic importance as primary parasitoids of aclerdids (Hemiptera: Aclerdidae) on grasses (Poaceae). We review the known species and describe three as new, bringing the total to eight. The species are characterised by means of a dichotomous key, diagnoses and illustrations and their known hosts are summarised.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:324E4AF7-3032-4573-98AC-CABAE316F33E     

A revision of the European species of Baeoneurella Dodd (Hymenoptera: Scelionidae)

ABSTRACT

The genus Baeoneurella Dodd, is resurrected, and Eumicrosoma Gahan, is treated as a junior synonym. A comparison between Baeoneurella and Telenomus Haliday, with emphasis on the T. floridanus group, is made to evaluate the generic limits of Baeoneurella, and synapomorphies for this genus are presented. The European species of Baeoneurella, including a new species, B. maceki Popovici, Masner and Polaszek, are described and illustrated, with new data concerning their distribution and habitats. Keys to the European species of this genus for females and for males are provided.

http://zoobank.org/urn:lsid:zoobank.org:pub:3F3ECAB2-FBF9-40BB-8447-06CEF1073038     

Review of the Chinese Bittacus (Mecoptera: Bittacidae) with descriptions of three new species

The current knowledge of the Chinese species of the hangingfly genus Bittacus Latreille, 1805 is reviewed. A checklist of the known species is provided. A key to species is compiled. Three species of Bittacus are described as new to science: Bittacus longantennatus sp. nov. from Sichuan and Hubei, Bittacus setigerus sp. nov. from Hubei and Guizhou, and Bittacus jiulianshanus sp. nov. from Jiangxi. http://www.zoobank.org/urn:lsid:zoobank.org:pub:5167E4E9-C123-4C1E-B33F-BCC7AF873BEA     

Two new species of Clinocentrus Haliday (Hymenoptera: Braconidae) from Argentina,with notes on South American species

The Argentine species of the genus Clinocentrus Haliday are revised. Two new species are described, Clinocentrus mamull sp. nov. and Clinocentrus roigi sp. nov., which represent the southernmost records of the genus in South America. Additionally Clinocentrus nigripectus Enderlein is redescribed and a key to known South American species is provided.     

Cryptic subarctic diversity: a new bumblebee species from the Yukon and Alaska (Hymenoptera: Apidae)

It is nearly 90 years since a bumblebee species has been described for the first time from Canada or the USA that was previously unrecognised by science. We describe and illustrate Bombus(Alpinobombus) kluanensis sp. nov. and begin to map its distribution. This species was discovered through analysis of Cytochrome c Oxidase subunit I barcodes and had been unsuspected on the basis of morphological and colour characteristics. So far, 247 female specimens considered to belong to this species are known from the subarctic mountains associated with the Denali Fault running from the Alaska Range (Alaska, USA) to Kluane in the Saint Elias Mountains (Yukon, Canada). A checklist of all species of the subgenus Alpinobombus worldwide is provided.

http://zoobank.org/urn:lsid:zoobank.org:pub:78D524DD-FFBA-4BC4-8D39-3EF4F6BDD39E     

A review of three species of Hesperonoe (Annelida: Polynoidae) in Asia,with descriptions of two new species and a new record of Hesperonoe hwanghaiensis from Korea

A taxonomic review of three Asian species of Hesperonoe (Annelida: Polynoidae) is presented. Hesperonoe hwanghaiensis Uschakov and Wu, 1959 Uschakov PV, Wu B. 1959. [The polychaete worms of the families Phyllodocidae and Aphroditidae (Polychaeta, errantia) from the Yellow Sea]. Arch Inst Oceanol Sinica. 1:1–40, pls. I–X. (In Chinese and Russian). [Google Scholar] (type locality: Qingdao, China) is redescribed based on five specimens collected from an intertidal site in the Korean coast of the Yellow Sea, as a new record of this species from Korea, which is the only known habitat outside the type locality. Hesperonoe coreensis sp. nov. is described based on nine specimens collected from two intertidal sites in the Korean coast of the Yellow Sea, including the same site as the habitat of H. hwanghaiensis. Hesperonoe japonensis sp. nov. is described by re-examination of many specimens collected from eight intertidal sites and a subtidal one in Japan, which was previously misidentified as H. hwanghaiensis. The three species are clearly distinguishable from one another by the species-specific morphology of the macrotubercles (or marked ridges) on the surface of elytra. All of the three species seemed to be commensal with the burrowing mud shrimp Upogebia major in intertidal flats, except for an additional probable host of another upogebid shrimp Austinogebia narutensis for H. japonensis sp. nov. in a subtidal habitat. The morphological characteristics and host species of the three Asian species are compared with all of the other five congeners known from the North American Pacific and Arctic Sea.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:51CB5C5B-0838-48AC-A0BA-8E63AA7628CB     

Notes on some palaearctic and oriental Systropha,with descriptions of new species and a key to the species (Hymenoptera: Apoidea: Halictidae)

An annotated list of the palaearctic and oriental species of the genus Systropha Illiger, 1806, includes additional records for some less well-known species. A lectotype is designated for Systropha hirsuta Spinola, 1839. Systropha christae Warncke, 1992, is synonymized with S. hirsuta. Systropha androsthenes and S. diacantha, spp. n., are described from the Gulf States. A key for the separation of the males is given.     

A taxonomic and zoogeographical census of the extant ant taxa (Hymenoptera: Formicidae)

The numbers of extant ant taxa in the world are analysed taxonomically and zoogeographically. A table detailing numbers of subfamilies, tribes and genera, and the numbers of species they contain, and one analysing the number of species per genus per zoogeographical region, are presented. Subfamilies are analysed by number and percentage of genera and species; the most speciose genera are isolated and listed in terms of absolute size and speciosity per subfamily. Tables detailing total numbers of genera and species per subfamily per region, and endemic genera per subfamily per region are given, and the endemicity and speciosity of genera are tabulated on a regional basis.     

An unusual,flagellum-bearing hydrobiid snail (Gastropoda: Rissooidea: Hydrobiidae) from Greece,with descriptions of a new genus and a new species

A new genus and a new species of Hydrobiidae, Agrafia wiktori sp. nov., from the Agrafa Mountains, northern Greece, are described based on the shell, protoconch, radula and male and female reproductive organs. In the protoconch sculpture and female reproductive organs A. wiktori resembles a typical member of Hydrobiidae Sadlerianinae, but the penis with tubular penial gland terminating within the flagellum and extending into the cephalic haemocoele has not been found in any other representative of Hydrobiidae. Instead, it is comparable with the penes and tubular penial glands of some non-hydrobiid Rissooidea (Bithyniidae, Bythinellidae, Amnicolidae, Emmericidae). Analysis of partial sequences of 18S ribosomal DNA and cytochrome oxidase subunit I mitochondrial DNA in A. wiktori place it in Hydrobiidae Sadlerianinae. The evolution of the bifid penis and tubular penial gland is discussed.     

Morphological and molecular separation between Macrocamptoptera grangeri Soyka and M. metotarsa (Girault) (Hymenoptera: Mymaridae)

ABSTRACT

Based on morphological and molecular evidence, the European fairyfly Herulia sundholmi Hedqvist syn. n. (Hymenoptera: Mymaridae) is removed from previous synonymy with the Nearctic Macrocamptoptera metotarsa (Girault), and synonymised with the Palaearctic M. grangeri Soyka, which is newly recorded from Georgia, Slovakia and Spain. Macrocamptoptera bulagarica (Donev) syn. n., known from Bulgaria and Italy, is also synonymised with M. grangeri. An updated key to females of the three currently recognised species of the rarely collected genus Macrocamptoptera Girault is given.     

Revision of Indian species of Baeus Haliday (Hymenoptera: Platygastroidea: Scelionidae)

ABSTRACT

Baeus senus Kozlov and Lê from Vietnam and B. primitus Rajmohana from India are the only two species of the genus known from the Oriental region. Thirty eight new species of Baeus are here described from India: B. acuminatus sp. n., B. agniparvathus sp. n., B. airavata sp. n., B. arachnophagus sp. n., B. bagheera sp. n., B. bharathiae sp. n., B. chakora sp. n., B. chitrasena sp. n., B. ciprianii sp. n., B. densipilosus sp. n., B. flaviscapus sp. n., B. gajakarna sp. n., B. giganteus sp. n., B. himalayanus sp. n., B. krishnareddyi sp. n., B. krumbiegeli sp. n., B. kubera sp. n., B. longiabdominalis sp. n., B. mahanetra sp. n., B. mareecha sp. n., B. menaka sp. n., B. nbairus sp. n., B. nicobarensis sp. n., B. pygmaeus sp. n., B. rachanae sp. n., B. rambha sp. n., B. ravana sp. n., B. reticulatus sp. n., B. sreedeviae sp. n., B. striatus sp. n., B. takshaka sp. n., B. tejaswii sp. n., B. tilottama sp. n., B. tripurasundari sp. n., B. tumburu sp. n., B. urvashi sp. n., B. vichitra sp. n. and B. xanthoclavatus sp. n. A key to the Indian species of Baeus is provided along with illustrations.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:878C5BFB-C2CB-4163-A08B-C761B4C6F087