On the fly fauna of the central region of the Kingdom of Saudi Arabia: new country records from Riyadh Region,with a list of associated fly species and zoogeographical remarks (Insecta: Diptera)

This study presents three genera and 10 species of flies (order: Diptera) as new country records from the Kingdom of Saudi Arabia (KSA). The flies were identified from Rawdhat Khorim National Park in Riyadh Region, the central region of KSA. The newly recorded genera are Glabellula (Mythicomyiidae), Phora (Phoridae) and Ceroptera (Sphaeroceridae), and the newly recorded species are Dilophus lingens (Bibionidae), Stichopogon deserti (Asilidae), Glabellula sp. (Mythicomyiidae), Phora sp. (Phoridae), Liriomyza lutea (Agromyzidae), Ceroptera aharonii (Sphaeroceridae), Trixoscelis deemingi, Trixoscelis migueli and Trixoscelis puncticornis (Trixoscelididae), and Physiphora leucotricha (Ulidiidae). Glabellula sp. and Phora sp. have been identified only to the genus level and are listed herein only because this is the first time they have been recorded in KSA, and additional specimens are required for their accurate identification to the species level. Also, this is the first record of Trixoscelis deemingi Wo?nica and Trixoscelis migueli Wo?nica from the Palaearctic Region. Distribution, diagnoses, remarks on biology and coloured photos are given for the recorded taxa. A list of associated fly species that were previously recorded in KSA and are identified from the study area in the present study is also presented. Zoogeographically, 33% of the identified fly species showed both Afrotropical and Palaearctic affinities, while 17% were of only Palaearctic affinity, and 8% were of only Afrotropical affinity. Elements from other regions were apparently nil. The results support the idea that the central Arabian Desert, which includes the present study area, is a transitional zone between the Palaearctic and Afrotropical regions.     

Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)

The South American Trogidae are revised. Three genera, Trox Fabricius, Omorgus Erichson (with two subgenera, Omorgus and Haroldomorgus Scholtz), and Polynoncus Burmeister, and 47 species are recorded from the continent and its islands. Trox is represented by the ubiquitous introduced species T. scaber (L.), Omorgus is represented by 13 species and Polynoncus by 33 species. Five new species are described and all others are re-described. A key to all taxa is provided. Adults of all species and their male genitalia are illustrated and their distributions mapped. Observations are presented on phylogeny, zoogeography and biology. The new species are O. capillaceus, O. indigenus, O. nocheles, P. erugatus and P. gibberosus. Four new synonymies are proposed (synonyms listed first): O. triestinae Pittino = P. suberosus (Fabricius); P. furcifer Pittino = P. bifurcatus (Vaurie); P. parafurcatus occidentalis Pittino and P. furcillifer Pittino = P. parafurcatus Pittino. Omorgus badeni (Harold) is reinstated as a valid species. Lectotypes are designated for O. batesi (Harold) and O. badeni (Harold).     

A description of Siphonophora portoricensis Brandt (Diplopoda: Siphonophorida: Siphonophoridae), with a Catalogue of Ordinal Representatives in the New World

Data on geographical distribution are provided for S. portoricensis Brandt, the type species of Siphonophora Brandt and the oldest species-group name in the genus, family, and Order; it appears to occur throughout Puerto Rico. It is characterized by fusion of the distal three podomeres of the anterior gonopods, which are sublinear in shorter, less mature males and bend caudad for varying lengths in longer individuals. In the New World, the Siphonophorida is represented by both families, the Siphonorhinidae by one genus and one species, Illacme plenipes Cook and Loomis, from San Benito County, California, and the Siphonophoridae by 10 genera, six monotypic, 66 extant nominal species, and one subspecies; two fossil species of Siphonophora have been proposed along with an unnamed fossil form of Siphonocybe Pocock. The following new combinations are officially recorded: Columbianum gracilicornis (Carl), Columbiozonium pearsei (Chamberlin), Cordillerium fuhrmanni (Carl), Yucatanium sabachana (Chamberlin), and Guatemalium barberi (Chamberlin). The Siphonophoridae ranges from southern California, northcentral Arizona, and central Texas to the Andes of central Peru and São Paulo state, Brazil. It also occurs on 10 islands in the Greater and Lesser Antilles, being newly recorded from Martinique, but apparently does not cross the Straits of Florida and is unknown from Florida, the Bahamas, and the Turks and Caicos Islands.     

Weevil (Coleoptera: Curculionoidea) seed-predators of Syzygium and Eugenia spp. (Myrtaceae), with description of the Alcidodes expansitarsis species group (Coleoptera: Curculionidae: Molytinae: Mecysolobini)

The weevils known to develop in Syzygium and Eugenia (Myrtaceae) fruit are reviewed. These weevils belong to the genera Curculio, Alcidodes, Acythopeus, Apotomorhinus (Curculionidae: Curculioninae, Molytinae and (the last two) Baridinae respectively) and Sitophilus (Dryophthoridae) in the Old World, and Atractomerus (Curculionidae: Anthonominae) in the Neotropics; two undescribed baridine species (one in Australia and the other in the USA) are also known to feed on the fruits, and the Omophorine Teleuropus (Curculionidae: Molytinae) has been found associated with them. Particular attention is paid to species of Alcidodes from aseasonal dipterocarp-dominated forests in South-East Asia. Two Bornean species (Alcidodes janetae sp. n. and A. eugeniophilus sp. n.) are the first of their genus to be reared from the fruit of Syzygium or Eugenia. These species, plus two other (A. expansitarsis sp. n. from Assam, Myanmar, Cambodia and Laos, and A. gymnasticus sp. n. from Malaysia) belong to a previously undescribed species group of Alcidodes; all four species are described and keyed.     

Revision of the southern African genus Frankenbergerius Balthasar with description of new taxa (Coleoptera: Scarabaeidae: Scarabaeinae)

The southern African genus Frankenbergerius Balthasar is revised. Two new species, F. opacus sp. n. and F. nitidus sp. n. (South Africa, Western Cape), and one new subspecies, F. armatus tuberculatus ssp. n. (South Africa, Mpumalanga), are described. F. imitativus (Péringuey) is considered a junior synonym of F. forcipatus (Harold). A key to the species and notes on biology are given.     

Mites of the family Oppiidae (Acari: Oribatida) from Turkey

Two new oppiid species (Gressittoppia luxtoni, Anomaloppia ozkani) and one new subspecies (Graptoppia sundensis acuta) are described from Turkish soils, and two known species are recorded for the first time. The genera Gressittoppia, Anomaloppia and Graptoppia are redefined and keys provided to the known species.     

Ant fauna (Hymenoptera: Formicidae) of the Socotra Archipelago (Yemen): zoogeography,distribution and description of a new species

The animal fauna of the Socotra Archipelago is influenced by three biogeographical regions, the Afrotropical, the Oriental and the Palaearctic. Consequently, the Archipelago shares faunal elements of these three regions in addition to unique endemic taxa. The ant fauna of Socotra Island was studied and is reviewed based on literature and newly collected material. In total, 28 species, belonging to 10 genera and four subfamilies, were collected from the main island. Eighteen of these (64%) are successful invasive species, seven are native (25%), and three are considered endemic (11%), Cardiocondyla longiceps Seifert, Monomorium elghazalyi sp. nov. and Monomorium nimihil Collingwood et al. Two genera are recorded for the first time from the island, Hypoponera Santschi, and Syllophopsis Santschi. Ten species are recorded for the first time, Cardiocondyla mauritanica Forel, Cardiocondyla minutior Forel, Monomorium atomum Forel, Monomorium dichroum Forel, Monomorium exiguum Forel, Pheidole pallidula (Nylander), Syllophopsis cryptobia (Santschi), Tetramorium pauper Forel, Tetramorium transformans Santschi and Hypoponera punctatissima (Roger). Ten invasive species are recorded from Socotra, reflecting human impacts on the Archipelago. These species are Tapinoma melanocephalum (Fabricius), Cardiocondyla emeryi Forel, Monomorium exiguum Forel, Pheidole indica Mayr, Syllophopsis cryptobia (Santschi), Tetramorium lanuginosum Mayr, Tetramorium simillimum (Smith), Tetramorium caldarium (Roger), Trichomyrmex destructor (Jerdon) and Trichomyrmex mayri (Forel). Our survey indicated a mixture of Afrotropical faunal elements (10 species, 36%), followed by cosmopolitan (nine species, 32%), Palaearctic (five species, 18%) and Oriental (four species, 14%) taxa. Two new synonyms of Monomorium exiguum Forel are proposed: Monomorium exiguum Forel = Monomorium baushare Collingwood & Agosti syn. nov. = Monomorium qarahe Collingwood & Agosti syn. nov. Tetramorium transformans Santschi is removed from synonymy with Tetramorium caldarium (Roger) and elevated to species rank. Ecological and biological notes for each species are given. Distribution maps for all species known from the Socotra Archipelago are provided.

http://zoobank/urn:lsid:zoobank.org:pub:89612083-9CE6-48E8-8975-1CE5334E098B     

Review of the genus Bolbonaso Emeljanov with checklist and key to Indian Caliscelidae (Hemiptera: Fulgoroidea)

A list of species of the family Caliscelidae Amyot et Serville known from India with data on distribution and sources for identification and a key to genera are given. The genus Bolbonaso Emeljanov, 2007 is revised. A new species, Bolbonaso chandri sp. nov., is described from Eastern India (Assam and Meghalaya States). Bolbonaso tapirifacies (Parshad, 1981) is redescribed and recorded for the first time from Southern India (Karnataka State). The lectotype is designated for Chirodisca eximia (Stål, 1859) which is recorded for the first time from Pakistan. New faunistic data in Nepal are listed for Delhina eurybrachydoides Distant, 1912.     

Checklist of Hydrophiloidea of Turkey (Coleoptera: Polyphaga)

The superfamily Hydrophiloidea of Turkey is catalogued. A total of 23 genera and 161 valid species and subspecies are recorded of which six species (3.7%) are recorded as endemic. Forty-eight species and two subspecies are from the family Helophoridae, three species are from the family Georissidae, seven species are from the family Hydrochidae, two species are from the family Spercheidae and 95 species and four subspecies are from the family Hydrophilidae. Distributions of these species within Turkey are given according to the 81 administrative divisions of the country. The catalogue is based on examination of the primary literature and also includes some records from specimens examined in collections. Taxa are arranged hierarchically under the categories of family, subfamily, tribe, subtribe (where recognized), genus, subgenus (where recognized), species and subspecies (where recognized).     

Report on the Raphitomidae Bellardi, 1875 (Mollusca: Gastropoda: Conoidea) from the China Seas

Twenty-seven Raphitomidae species belonging to nine genera, are recorded from the China Seas, including two new species, which are described here: Asperdaphne paramoretonica sp. nov., Daphnella inangulata sp. nov. Eight species are recorded for the first time from the China Seas.     

A revision and phylogenetic re-evaluation of the family Schizopodidae (Coleoptera,Buprestoidea)

The Schizopodidae (formerly subfamily Schizopodinae, family Buprestidae) is elevated to family status, based on a review of its morphology including wing venation, male and female genitalia, larvae, and also a cladistic analysis of in-group and out-group comparisions. Two tribes are recognized: Schizopodini LeConte with one genus, Schizopus LeConte and Dystaxiini Théry with two genera: Dystaxia LeConte and Glyptoscelimorpha Horn. The genus Schizopus contains two species: S. laetus LeConte and S. sallei Horn with one new subspecies, S. sallei nigricans Nelson. The genus Dystaxia includes two species: D. murrayi LeConte, and D. elegans Fall. The genus Glyptoscelimorpha is separated into two subgenera: G. (s. str.) with two species, G. marmorata Horn, and G. viridis Chamberlin; and G. (Dystaxiella) Knull, new status, with one species, G. (D.) juniperae (Knull), new combination, with a new subspecies, G. (D.) juniperae viridiceps Nelson. All taxa are described and information on variation, type localities, distribution, biology, and comparisons are given for each species. Keys to taxa for tribes to subspecies are provided. Illustrations for all species, including habitus, male and female genitalia and other parts, as well as cladograms of relationships are presented. Lectotypes are designated for the following: Yermoella helferi Obenberger, Dystaxia elegans Fall and Glyptoscelimorpha marmorata Horn. Tables of taxa, characters and character states used in the cladistic analysis are included.     

Biodiversity of the scentless plant bugs (Hemiptera: Rhopalidae) in southern South America

This paper provides a comprehensive faunal survey of the Rhopalidae from southern South America with emphasis on the fauna of Argentina, based on published and unpublished data. The biodiversity from Argentina comprises 38 recorded species from six genera: Arhyssus Stål 1870 (one species), Liorhyssus Stål 1870 (two species), Niesthrea Spinola 1837 (five species), Xenogenus Berg 1883 (two species), Harmostes Burmeister 1835 (20 species), and Jadera Stål 1862 (eight species). We established accurate distributions for most of the species and report new distributional information for 27 of them. Most of these records are from Argentinean provinces, but we also report five new country records: three from Argentina – Harmostes (Harmostes) splendens Harris 1944, H. (Neoharmostes) bergi Göllner-Scheiding 1998 and Arhyssus tricostatus (Spinola 1852); one from Brazil – Xenogenus picturatum Berg 1883; and one from Paraguay – Harmostes (Harmostes) gravidator (Fabricius 1794).     

Rediscovery of the Hawaiian endemic bark beetle Xyleborus pleiades Samuleson, 1981 on Moloka‘i,with records of three new exotic bark beetles for the island (Coleoptera: Curculionidae: Scolytinae: Xyleborini)

ABSTRACT

During the course of a targeted bark beetle survey of the Hawaiian Islands, insect sampling undertaken in native forest biotopes on Moloka‘i island revealed the presence of one native and three exotic species of xyleborine bark beetles (Coleoptera: Curculionidae: Scolytinae: Xyleborini) hitherto not recorded from the island. We present detailed new island records for these species, which include the second record of Xyleborus pleiades Samuelson, 1981, an endemic Hawaiian species not recorded in over 50 years, and previously known only from two specimens from the island of Maui. The three newly recorded introduced species are widespread pantropical beetles of actual or potential pest status: Xyleborus affinis Eichhoff, 1867, Xyleborinus saxesenii (Ratzeburg, 1837), and Xylosandrus crassiusculus (Motschulsky, 1866). The importance of the records is briefly discussed, and all newly recorded species are illustrated in colour photographs.     

A preliminary account of the fly fauna in Garf Raydah Nature Reserve,Kingdom of Saudi Arabia,with new records and biogeographical remarks (Diptera: Insecta)

A preliminary list of Diptera (Insecta) in Garf Raydah Nature Reserve (GRNR), Kingdom of Saudi Arabia (KSA) with remarks on their zoogeographic affinities is presented. This is the first in a series of planned faunal studies on different insect orders as an output of a project proposed to study the entire entomofauna of GRNR. A total number of 177 Diptera species belonging to 130 genera, representing 38 families has been listed. Some species have been identified only to genus and listed herein as genera that were not previously recorded from KSA or of a taxonomic or faunal importance. The list records five families, 11 genera and 31 species for the first time in KSA, namely: the families Acroceridae, Asteiidae, Chamaemyiidae, Heleomyzidae and Pipunculidae; the genera Acrocera (Acroceridae), Eremisca (Asilidae), Asteia (Asteiidae), Chamaemyia sp. (Chamaemyiidae), Ochthera (Ephydridae), Physoconops (Conopidae), Pseudoleria (Heleo-myzidae), Salentia (Therevidae), Sapromyza (Lauxaniidae), Sternobrithes (Stratiomyidae), Tomosvaryella (Pipunculidae); and the species Apoclea algira (Linnaeus), Apoclea femoralis (Wiedemann), Promachus rectangularis Loew, Eremisca sp. and Oligopogon nitidus Efflatoun [Asilidae]; Phycus sp., Salentia sp. and Thereva nobilitata (Fabricius) [Therevidae]; Drapetis flavipes Macquartand Platypalpus flavicornis (Meigen) [Hybotidae]; Acrocera sp.1 and Acrocera sp.2 [Acroceridae]; Aspida-cantha atra Kertesz and Sternobrithes sp. [Stratiomyidae]; Tomos-varyella sp.1 and Tomosvaryella sp.2 [Pipunculidae]; Eristalinus taeniops (Wiedemann) and Eumerus amoenus Loew [Syrphidae]; Asteia sp. [Asteiidae]; Chamaemyia sp. [Chamaemyiidae]; Physoconops sp., and Thecophora fulvipes Rubineau-Desvoidy [Conopidae]; Ochthera sp. [Ephydridae]; Pseudoleria pectinata (Loew) [Heleomyzidae]; Cestrotus megacephalus Loew and Sapromyza sp. [Lauxaniidae]; Campylocera oculata Hendel [Pyrgotidae]; Australosepsis niveipennis (Becker) [Sepsidae]; Metasphenisca transilis Munro [Tephritidae]; Anthomyia xanthopus (Hennig) [Anthomyiidae]; and Sarcophaga inaequalis Austen [Sarcophagidae]. More than 50% of the recorded fly species are of an Afrotropical affinity, whereas the Palaearctic elements did not exceed 20%. Only 7% of the species were of both Afrotropical and Palaearctic affinities, and Oriental elements were close to nil. This emphasizes the fact that the southwestern part of KSA, including the study area, belonging to Asir Province, ought to be part of the Afrotropical Region rather than the the Palaearctic Region.     

Report on Crassispirinae Morrison, 1966 (Mollusca: Neogastropoda: Turridae) from the China Seas

The present paper reports 32 turrid species from the China Seas, belonging to eight genera of the subfamily Crassispirinae of the family Turridae. Four new species are described: Funa cretea sp. nov., Inquisitor plurivaricis sp. nov., Inquisitor vividus sp. nov. and Ptychobela resticula sp. nov. Eight species are recorded for the first time from the China Seas.     

Apelaunothripini from the Philippines (Thysanoptera: Phlaeothripidae)

Summary

The relationships of the phlaeothripine tribe Apelaunothripini is discussed. Four genera and thirteen species are recorded from the Philippines, including the following two new genera and nine new species: Apelaunothrips cephalicus sp. nov., A. fasciatus sp. nov., A. flavicornis sp. nov., A. marginalis sp. nov., A. philippinensis sp. nov., A. rostratus sp. nov., A. simpliceps sp. nov., Lizalothrips luzonensis gen. and sp. nov., Paradexiothrips bispinosus gen. and sp. nov. Keys are provided to the four genera included in the Apelaunothripini, and to the ten Apelaunothrips species from the Philippines.     

Australian thrips of the Haplothrips lineage (Insecta: Thysanoptera)

Keys are presented to the 11 genera and 50 species, including 21 new species, of Thysanoptera in Australia that are related to the worldwide genus Haplothrips Amyot and Serville. These taxa belong to what, in recent literature, has been called the “Haplothrips‐lineage”, that is, one of the three major radiations among the 2700 species and 350 genera of Thysanoptera Phlaeothripinae. The group is redefined, and the available tribal name Haplothripini shown to be appropriate. The character states on which the definition is based are discussed, and a list given of the 34 genera worldwide that can be included. The Australian species in these genera exhibit a diversity of biologies. Three genera involve species that invade galls induced by other thrips: Androthrips monsterae (Moulton) from New Guinea is newly recorded from Australia; Mesothrips jordani Zimmermann from South‐East Asia is newly recorded from Australia, with two new synonyms; the Asia‐Pacific genus, Euoplothrips Hood, includes two species in northern Australia. Three haplothripine genera are associated with grasses: one widespread genus, Apterygothrips Priesner, considered polyphyletic, includes only one Australian species; the only known species of Dyothrips Kudo is Oriental but extends into tropical Australia; Podothrips Hood, a circum‐tropical genus of thrips predatory on grass‐living coccoids, has 11 Australian species, six newly described. One grass‐associated genus, Bamboosiella Ananthakrishnan, is excluded from the Haplothripini. An Oriental genus of leaf‐ and flower‐living species, Dolichothrips Karny, includes one species in northern Australia, D. reuteri (Karny); Membrothrips Bhatti in which this species has been placed is considered a synonym. Karnyothrips Watson includes two species introduced to Australia, both predators of coccoids. Priesneria Bagnall includes three species from Australia, of which one fungus‐feeding species is newly described. Xylaplothrips Priesner is a widespread but ill‐defined, polyphyletic genus that currently includes three little‐known Australian species. Haplothrips is the main focus of this study, and character state variation among the Australian species is discussed. In total 24 species of Haplothrips from Australia are recognised, 14 being new species. Also included are the following, one new generic synonym, four new species synonyms, and one new combination. The 250 species worldwide in the genus are usually associated with the flowers of Asteraceae and Poaceae. In Australia, several of the species are specific to flowers, particularly of Poaceae and Cyperaceae but not Asteraceae. Almost half of these Australian species are presumed to be predatory on other small arthropods, and two have unusual host associations—with salt marsh Chenopodiaceae, and with the sori of Dicksonia tree ferns.     

A revision of the subtribe Hippomelanina,part II: Gyascutus (Gyascutus) LeConte (Coleoptera: Buprestidae)

The subgenus Gyascutus (Gyascutus) is revised and 12 species are recognized: G. planicosta (LeConte) with three subspecies— G. p. planicosta (LeConte), stat. nov.; G. p. obliteratus (LeConte), stat. nov., with G. pistorius Casey as a syn. nov.; and G. p. cribriceps Casey, stat. nov.; with syn. nov. as follows: G. acutangulus Casey, G. amplus Casey, G. amplus aerata Casey, G. egregius Casey, and Hippomelas (G.) lariversi Barr; G. insularis (Helfer), comb. nov.; G. castaneus (Helfer), comb. nov.; G. fulgidus (Barr), comb. nov.; G. pacificus (Chamberlin), comb. nov.; G. paragranulatus Nelson, sp. nov.; G. granulatus (Van Dyke), comb. nov.; G. westcotti Nelson, sp. nov.; G. carolinensis Horn with syn. nov. as follows: G. juniperinus Wickham, G. obesus Casey, G. obesus aeneoviridis Casey, G. compactus Casey, G. compactus solidus Casey, G. cylindrinus Casey, G. debilis Casey, and G. tenuis Casey; G. dianae (Helfer), comb. nov.; G. allenrolfeae (Verity), comb. nov.; and G. jeanae (Nelson), comb. nov. All species and subspecies are described and information on type localities, variation, distribution, biology, and comparisons is presented. Keys to the species and subspecies are provided. Photographs of each species and drawings of body parts, including male genitalia, are included.     

The species of four genera of Metopiinae (Hymenoptera: Ichneumonidae) in Britain,with new host records and descriptions of four new species

Two genera of Metopiinae are recorded for the first time from the British Isles, Ischyrocnemis Holmgren and Synosis Townes. An account is also given of a further two genera, Apolophus Townes and Stethoncus Townes, that have been recently recorded from Britain but remain little known. Apolophus and Synosis are shown to be koinobiont endoparasitoids of Lepidoptera larvae of the families Schreckensteiniidae and Yponomeutidae, respectively. Four species are described as new: Stethoncus monopicida sp. nov., Synosis caesiellae sp. nov., Synosis fieldi sp. nov., and Synosis parenthesellae sp. nov. A key to the Western Palaearctic species of Synosis is provided.