A new species of Nemedina Chandler from mid-Asia (Diptera: Empidoidea) resolves the phylogenetic position of this enigmatic genus

Nemedina zaitsevi sp. nov. is described from Kazakhstan and includes the first extant male specimens of this genus. The combination of male genitalic features (e.g. long and narrow gonocoxal apodemes, shortened hypandrium), absence of female tergite 10, and absence of epipharyngeal blades strongly support the hypothesis that Nemedina Chandler and the Atelestinae are sister groups.