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Birds are unique among living vertebrates in possessing pneumaticity of the postcranial skeleton, with invasion of bone by the pulmonary air-sac system. The avian respiratory system includes high-compliance air sacs that ventilate a dorsally fixed, non-expanding parabronchial lung. Caudally positioned abdominal and thoracic air sacs are critical components of the avian aspiration pump, facilitating flow-through ventilation of the lung and near-constant airflow during both inspiration and expiration, highlighting a design optimized for efficient gas exchange. Postcranial skeletal pneumaticity has also been reported in numerous extinct archosaurs including non-avian theropod dinosaurs and Archaeopteryx. However, the relationship between osseous pneumaticity and the evolution of the avian respiratory apparatus has long remained ambiguous. Here we report, on the basis of a comparative analysis of region-specific pneumaticity with extant birds, evidence for cervical and abdominal air-sac systems in non-avian theropods, along with thoracic skeletal prerequisites of an avian-style aspiration pump. The early acquisition of this system among theropods is demonstrated by examination of an exceptional new specimen of Majungatholus atopus, documenting these features in a taxon only distantly related to birds. Taken together, these specializations imply the existence of the basic avian pulmonary Bauplan in basal neotheropods, indicating that flow-through ventilation of the lung is not restricted to birds but is probably a general theropod characteristic. 相似文献
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A 629RKLKK633 motif in the hinge region controls the androgen receptor at multiple levels 总被引:1,自引:1,他引:0
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This paper uses recent advances in time-series modeling to derive long-horizon forecasts of commodity price volatility which incorporate investors' expectations of volatility. Our results are promising. We compare several different forecasts of commodity price volatility, which we divide into three categories: (1) forecasts using only expectations derived from options prices; (2) forecasts using only time-series modeling; and (3) forecasts which combine market expectations and time-series methods. The forecasts in (1) and (2) are used extensively in the literature, while those in (3) are new in this paper. On comparing these different forecasts, we find that our proposed forecasts from category (3) outperform both market expectations forecasts and time-series forecasts. This result holds both in and out of sample for virtually all commodities considered. 相似文献
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