New fossil ommatid (Coleoptera: Archostemata: Ommatidae) from Jehol Biota of western Liaoning, China

In this paper one new genus and one new species of fossil ommatid, Odontomma trachylaena gen. et sp. nov., is described and illustrated from the Late Jurassic or Early Cretaceous Yixian Formation of western Liaoning, China. Based on wide epipleural space of elytron with several rows of cells, the new taxa is attributed to tribe Brochocoleini of Ommatidae and compared with genus Brochocoleus. Due to the fact that most other Chinese Mesozoic ommatids are not complete, the complete preservation of the new specimens reported here enabled us to determine the characteristics of the body of these ancient ommatid. Moreover, based on the well-preserved individual of this new genus, we may suppose the affinities of the new species with extant and extinct groups.     

A new fossil genus of Mesosciophilidae （Diptera, Nematocera） with two new species from the Middle Jurassic of Inner Mongolia, China

A new genus within the family Mesosciophilidae,Jurasciophila gen.nov.,with two new species,J.curvula gen.et sp.nov.and J.lepida gen.et sp.nov.,are described and illustrated.They are established based on fossil specimens with bodies and complete wings.All of them were collected from the late Middle Jurassic Jiulongshan Formation of Daohugou in southeastern Inner Mongolia,China.Comparing with two known genera,Mesosciophilina (Kovalev,1985) and Mesosciophilopsis (Blagoderov,1994),the new genus has transitional characters.The distribution pattern of Mesosciophilidae is briefly discussed.     

Two new ornithurine birds from the Early Cretaceous of western Liaoning, China

We describe two new ornithurine birds from the Early Cretaceous Jiufotang Formation of western Liaoning, northeast China: Yanornis martini gen. et sp. nov. and Yixianornis grabaui gen. et sp. nov. They represent the best fossil record of ornithurine birds known from the Early Cretaceous. They are more advanced than the most primitive ornithurine Liaoningornis, and are more similar to the other two Chinese Early Cretaceous ornithurines Chaoyangia and Songlingornis. Compared with Confuciusornis, Liaoxiornis and Eoenantiornis from the same age, the two new birds show remarkable advanced characteristics and suggest the presence of powerful flight capability like modern birds. Compared with Yixianornis and Chaoyangia, Yanornis is larger, with a more elongated skull and relatively long wings. The new discoveries indicate that by the Early Cretaceous both enantiornithine and ornithurine birds had already radiated significantly. The flight structures of Yanornis and Yixianornis are hardly distinguishable from those of modern birds; however, both retain a few primitive traits such as teeth on the jaws, wing claws and pubic symphysis, which exclude them from being the most recent ancestor of all extant birds.     

A new species from an ornithuromorph（Aves： Ornithothoraces）dominated locality of the Jehol Biota

We report on a new species of ornithuromorph bird, Iteravis huchzermeyeri gen. et sp. nov., from the previously unreported Sihedang locality of the Lower Cretaceous Yixian Formation, the oldest ornithuromorph bearing deposit in the world. Unlike most other Cretaceous localities, specimens from this new quarry are largely referable to Ornithuromorpha, similar to the Lower Cretaceous Aptian Xiagou Formation in Gansu Province. Also similar to the Xiagou avifauna, the fauna at Sihedang is largely dominated by a single taxon（described here）.Differences in faunal dominance may suggest the Sihedang records a unique ecological habitat. This may also explain the dominance of Gansus in the younger Xiagou Formation locality and suggests that previous hypotheses regarding the shift in dominance between Enantiornithes and Ornithuromorpha need to be reassessed in terms of potential ecological biases due to limited sampling. Furthermore, the recognition of an ornithuromorph dominated locality in the Sihedang significantly weakens the signal of such an inferred trend. Compared to most Jehol birds, the new specimen is relatively better preserved in three dimensions revealing morphological details of the skeleton, as well as preserves feather impressions including a rectricial morphology previously unknown among Mesozoic birds.     

A new family of primitive mammal from the Mesozoic of western Liaoning, China

A fairly well-preserved specimen of a new primitive mammal, Repenomamus robustus gen. et sp. nov., has been described from the Yixian Formation (Lower Cretaceous), western Liaoning, China. This animal has several derived characteristics, such as well-developed den-tary/squamosal articulation, reduced number of tooth, differentiation of premolars and molars in postcanine teeth, presence of a dorsal process of the premaxilla that is not in contact with the nasal, closed medial wall of the orbit, and presence of fingerlike promontorium on the petrosal. It also retains some primitive reptile-like features. It is the most primitive taxon among the three mammals known from the Jehol Biota, and also represents the largest mammal of Mesozoic age over the world.     

New dragonflies （Insecta： Odonata： Gomphaeschnidae） from the Yixian Formation in Inner Mongolia,China

Two fossil dragonflies from the Upper Jurassic to Lower Cretaceous Yixian Formation in Liutiaogou Village, Ningcheng County, Inner Mongolia, China are described and illustrated. They are assigned to two new genera and species, i.e., Sophoaeschna frigida gen. et sp. nov. and Falsisophoaeschna generalis gen. et sp. nov. within the family Gomphaeschnidae Tillyard ＆ Fraser, 1940. This is the first report of Odonata from Yixian Formation in Inner Mongolia and the second record of fossil Gomphaeschnidae from China.     

New dragonflies (Insecta: Odonata: Gomphaeschnidae) from the Yixian Formation in Inner Mongolia, China

Two fossil dragonflies from the Upper Jurassic to Lower Cretaceous Yixian Formation in Liutiaogou Village, Ningcheng County, Inner Mongolia, China are described and illustrated. They are assigned to two new genera and species, i.e., Sophoaeschna frigida gen. et sp. nov. and Falsisophoaeschna generalis gen. et sp. nov. within the family Gomphaeschnidae Tillyard & Fraser, 1940. This is the first report of Odonata from Yixian Formation in Inner Mongolia and the second record of fossil Gomphaeschnidae from China.     

A new Early Cretaceous lizard with well-preserved scale impressions from western Liaoning, China

A new small lizard, Liaoningolacerta brevirostra gen. et sp. nov., from the Early Cretaceous Yixian Formation of western Liaoning is described in detail. The new specimen was preserved not only by the skeleton, but also by the exceptionally clear scale impressions. This lizard can be included within the taxon Scleroglossa based on its 26 or more presacrals, cruciform interclavicle with a large anterior process, moderately elongated pubis, and slightly notched distal end of tibia. The scales vary evidently in size and shape at different parts of body: small and rhomboid ventral scales, tiny and round limb scales, and large and longitudinally rectangular caudal scales that constitute the caudal whorls. This new finding provides us with more information on the lepidosis of the Mesozoic lizards.     

A new iguanodontid (Jinzhousaurus yangi gen. et sp. nov.) from the Yixian Formation of western Liaoning, China

A new iguanodontid,Jinzhousaurus yangi gen. et sp. nov., is erected based on the cranial and dental morphology of a specimen from the Yixian Formation of western Liaoning. Although a few primitive characteristics remain,Jinzhousaurus yangi display a number of characteristics similar to those seen in derived iguanodontids. Other interesting characteristics include the absence of antorbital fenestra and the frontal excluded from the participation in orbit, which are closer to the condition in hadrosaurids. The unusual combination of the characteristics provides important data for assessing morphological evolution toward hadrosaurs. The new discovery also represents the first reported large-sized dinosaur from Jehol fauna in western Liaoning, and increases the diversity of this famous fauna. Furthermore, the discovery of a derived iguanodontid from the Yixian Formation provides further evidence supporting a Cretaceous age of the Jehol fauna, which is consistent with an isotopic age.     

A long-tailed,seed-eating bird from the Early Cretaceous of China

The lacustrine deposits of the Yixian and Jiufotang Formations in the Early Cretaceous Jehol Group in the western Liaoning area of northeast China are well known for preserving feathered dinosaurs, primitive birds and mammals. Here we report a large basal bird, Jeholornis prima gen. et sp. nov., from the Jiufotang Formation. This bird is distinctively different from other known birds of the Early Cretaceous period in retaining a long skeletal tail with unexpected elongated prezygopophyses and chevrons, resembling that of dromaeosaurids, providing a further link between birds and non-avian theropods. Despite its basal position in early avian evolution, the advanced features of the pectoral girdle and the carpal trochlea of the carpometacarpus of Jeholornis indicate the capability of powerful flight. The dozens of beautifully preserved ovules of unknown plant taxa in the stomach represents direct evidence for seed-eating adaptation in birds of the Mesozoic era.     

Discovery of a pterodactylid pterosaur from the Yixian Formation of western Liaoning, China

A well-preserved pterosaur with nearly complete skull is described from the Lower Cretaceous Yixian Formation at Sihetun in western Liaoning. It is characterized by a low and long crestless skull, slender and pointed teeth, long metacarpal, nearly equal length of metatarsals I–III and short pedal digit V. It is referred to a new genus and species of the family Pterodactylidae:Haopterus gracilis gen. et sp. nov. This is the first pterosaur with a nearly complete skull from the Jehol Biota; it also represents the first non-controversial fossil record of Pterodactylidae in Asia.Haopterus is more derived thanPterodactylus from the Late Jurassic Solnhofen in Germany. This discovery extends the distribution of the family Pterodactylidae from Europe and Africa to Asia and its latest occurrence from the Late Jurassic to the Early Cretaceous. The discovery ofHaopterus gracilis provides further evidence for the study of the origin and radiation of the Jehol Biota; it also sheds new light on the evolution and distribution of pterosaurs in the late Mesozoic.     

京西早白垩世丸甲（昆虫纲，鞘翅目）化石

根据北京西山下白垩统卢尚坟组的甲虫化石，建立化石丸甲二新属三新种：Ｍｅｓｏｂｙｒｒｈｕｓｔａｎａｅｇｅｎ．ｅｔｓｐ．ｎｏｖ，Ｍｅｓｏｂｙｒｒｈｕｓｐａｒｖｕｓｇｅｎ．ｅｔｎｏｖ．和Ｆａｎｇｓｈａｎｅｌａｓｔｏｌｉｄａｇｅｎ．ｅｔｓｐ．ｎｏｖ．．初步探讨了丸甲科昆虫的演化，认为丸甲科是鞘翅目中很古老的一类，和其它科关系不大，自中生代后期以来演化十分缓慢     

New discoveries from the classic Quaternary mammalian fossil area of Yanjinggou, Chongqing, and their chronological explanations

Yanjinggou in Wanzhou District of Chongqing is one of the earliest reported and most famous Quaternary mammalian fossil areas in China. The fauna from this area used to be taken as a benchmark for the biochronological comparison of Quaternary paleontology of South China, but the chronology of this fauna has many controversies for the lack of exact locational and stratigraphical records. The present article, on the basis of recent investigations and discoveries, discusses and explains the distribution and biochronology of the mammalian faunas in the Yanjinggou area. The newly discovered Dayakou fissure fauna, including Rhizomys troglodytes, Homotherium sp., Panthera pardus, Stegodon orientalis, Dicerorhinus sumatrensis, Hesperotherium sinense, Sus sp., Cervavitus fenqii, Cervus sp. and Muntiacus sp., is correlated to the middle Early Pleistocene. The presence of Early Pleistocene mammalian fauna in the Yanjinggou area is therefore confirmed. The “Wanhsien fauna” or “Yenchingkou (=Yanjinggou) fauna” as a Mid-Late Pleistocene biostratigraphical datum should be abandoned. The Dayakou fauna and the Upper Cave fauna of Pingba, both in the Yanjinggou area, are correlated to the middle Early Pleistocene and the early Middle Pleistocene in age, respectively.     

New fossil elaterids （Insect： Coleoptera： Polyphaga： Elateridae） from the Middle Jurassic of Inner Mongolia, China

A new genus with a new species of fossil elaterid, Paraprotagrypnus superbus gen. et sp. nov., from the Middle Jurassic Jiulongshan Formation in Daohugou Village (N41°18.979′, E119°14.318′), Shantou Township, Ningcheng County, Inner Mongolia, China is described and illustrated. The genus Paraprotagrypnus belongs to the family of Elateridae, subfamily of Protagrypninae and tribe of Protagrypnini. Some primitive characters of the new genus and new species shed light on the ages between the Jiulongshan Formation in Inner Mongolia and the Yixian Formation in the western Liaoning Province of China. The habitat of the new species is briefly discussed.     

Diverse transitional giant fleas from the Mesozoic era of China

Fleas are one of the major lineages of ectoparasitic insects and are now highly specialized for feeding on the blood of birds or mammals. This has isolated them among holometabolan insect orders, although they derive from the Antliophora (scorpionflies and true flies). Like most ectoparasitic lineages, their fossil record is meagre and confined to Cenozoic-era representatives of modern families, so that we lack evidence of the origins of fleas in the Mesozoic era. The origins of the first recognized Cretaceous stem-group flea, Tarwinia, remains highly controversial. Here we report fossils of the oldest definitive fleas--giant forms from the Middle Jurassic and Early Cretaceous periods of China. They exhibit many defining features of fleas but retain primitive traits such as non-jumping hindlegs. More importantly, all have stout and elongate sucking siphons for piercing the hides of their hosts, implying that these fleas may be rooted among the pollinating 'long siphonate' scorpionflies of the Mesozoic. Their special morphology suggests that their earliest hosts were hairy or feathered 'reptilians', and that they radiated to mammalian and bird hosts later in the Cenozoic.     

中国凹颜叶蜂族研究(膜翅目:叶蜂科)(英文)

本文记述中国凹颜叶蜂亚科凹颜叶蜂族一新属和三新种：细足叶蜂属Anheterusgen．nov．新屋,细足叶蜂Anheterustenuisp．nov．新种,郑氏细足叶蜂Anheteruszhengisp．nov．新种,长齿细足叶蜂Anheterusjangngshanussp．nov．新种。同时报道了中国叶峰科一新记录属科,黑鳞凹颜叶蜂HeterarthruskamtchaticaMalaise．     

中国潜叶蜂族新属种(膜翅目:叶蜂科)(英文)

描述了中国潜叶蜂族两新属三新种：短角额潜叶蜂Sinoscoliabrevicortnisgen.etsp.now.新属新种,故背鞘潜叶蝶Paraparnarubiginosagenetsp.now.新属材种,郑氏臀潜叶峰Afusazhengisp.nov.新种。指出臀游计蜂属是中国新记录属.     

Large Cretaceous sphenodontian from Patagonia provides insight into lepidosaur evolution in Gondwana

Sphenodontian reptiles successfully radiated during Triassic and Jurassic times, but were driven almost to extinction during the Cretaceous period. The sparse Early Cretaceous record of sphenodontians has been interpreted as reflecting the decline of the group in favour of lizards, their suspected ecological successors. However, recent discoveries in Late Cretaceous beds in Patagonia partially modify this interpretation. Numerous skeletons of a new sphenodontian, Priosphenodon avelasi gen. et sp. nov., were collected from a single locality in the Cenomanian-Turonian Candeleros Formation, where it is more abundant than any other tetrapod group recorded in the quarry (for example, Crocodyliformes, Serpentes, Dinosauria and Mammalia). Adult specimens of Priosphenodon reached one metre in length, larger than any previously known terrestrial sphenodontian. Here we propose, using available evidence, that sphenodontians were not a minor component of the Cretaceous terrestrial ecosystems of South America, and that their ecological replacement by squamates was delayed until the early Tertiary. The new discovery helps to bridge the considerable gap in the fossil record (around 120 million years) that separates the Early Cretaceous sphenodontians from their living relatives (Sphenodon).     

First ceratopsid dinosaur from China and its biogeographical implications

Ceratopsid dinosaurs represent one of the best known dinosaur groups in the Late Cretaceous, and their unquestionable fossil re-cords are exclusively restricted to western North America.Here we report a new ceratopsid dinosaur, Sinoceratops zhuchengensis gen.et sp.nov., from the Upper Cretaceous Wangshi Group of Zhucheng, Shandong Province, China.Cladistic analysis places this new taxon as the only known ceratopsid from outside North America, in a basal position within the Centrosaurinae.It is con-siderably...