Poisonous plants of Utah

Bird species density, diversity, and species richness in relation to habitat and seasonal variations were studied in the Betatakin Canyon area of Navajo National Monument, Arizona. The two most prominent habitat types are a riparian forest deep in the canyon bottom and a mature pinyon-juniper woodland on the mesas and slopes above the canyon. One hundred thirty-five species of birds were encountered during the study and recorded by season and habitat. The avifauna assemblages demonstrate definite habitat selection into groups associated with the riparian and pinyon-juniper woodland communities. Diversity, density, and species richness were greatest in the riparian habitat during the spring and summer months. The differences in bird community composition were greater between seasons than between habitats. Pinyon pine and Gambel oak were highly selected as perch sites in the pinyon-juniper and riparian areas, respectively.     

Habitat of three rare species of small mammals in juniper woodlands of southwestern Wyoming

Southwestern Wyoming constitutes the northern limit of the ranges of the cliff chipmunk ( Tamias dorsalis ), pinyon mouse ( Peromyscus truei ), and canyon mouse ( P. crinitus ). In addition to trying to determine their presence in the region, we wanted to identify habitat characteristics commonly used by each of these species. We used Sherman live-traps to sample 14 sites representing 2 distinct habitat types in 1998 and 1999: juniper-rocky slopes and juniper cliffs. Seventeen habitat characteristics were measured at capture locations for each species and compared with randomly located points. Best subsets multiple logistic regression was used to construct models that distinguish between used and available habitat for each species. The cliff chipmunk occurred in both rocky slopes and cliffs. The pinyon mouse was also captured in rocky slopes and cliffs and was most often captured in locations in the interior of the juniper woodland with high tree canopy cover, high forb cover, and low density of rock outcrops. The canyon mouse was captured only in cliffs at sites consisting of high forb cover, high rock cover, and high tree density.     

Effects of habitat type and degradation on avian species richness in Great Basin riparian habitats

The overwhelming majority of bird species in the Great Basin region are found in riparian habitats. However, most previous research on the impact of change in habitat condition through degradation on these bird communities failed to account for the large intersite differences, in both habitat type and extent of degradation. We examined songbird communities in 4 riparian habitat types (meadows, willow-birch-, and aspen-dominated forest stands) during summers 1994 (last year of a 7-yr drought) and 1995 (following the 6th wettest winter recorded) in the Toiyabe Mountain Range of central Nevada. Habitat degradation significantly influenced bird species richness in riparian areas, but the impact was dependent upon habitat type. While meadow bird communities were affected adversely by habitat degradation, with significant drops in species richness on degraded sites, bird species richness in forested riparian habitats was consistently greater on degraded sites. Data for the 6 most common species seen during our study indicated that degradation may have influenced distribution of American Robins ( Turdus migratorius ) and Yellow Warblers ( Dendroica petechia ), but habitat type was the best predictor of abundance for House Wrens ( Troglodytes aedon ), Red-naped Sapsuckers ( Sphyrapicus nuchalis ), Warbling Vircos ( Vireo gilvus ), and Brewer's Blackbirds ( Euphagus cyanocephalus ). Avian species diversity in meadow habitats may be linked to moisture levels during specific times of the year. Diversity increased during the pre-migratory period of the dry year (1994) when compared with that of the breeding season, but was unchanged in the wet year (1995).     

Variation in ant populations with elevation, tree cover, and fire in a pinyon-juniper–dominated watershed

Climate change and fire suppression have facilitated expansion of pinyon-juniper woodlands into sagebrush-steppe ecosystems of the Great Basin, USA, resulting in a loss of biological diversity. To assess the effects of using prescribed fire in restoration efforts, ant abundance, species richness, and composition were examined pre- and post-burn along the elevation and tree cover gradients encompassed by a pinyon-juniper woodland in a central Nevada watershed. Ants were sampled using pitfall traps in 6 sites for the elevation study and in 2 sites for the tree cover study, representing paired burn and control sites in a randomized block design. Vegetation and ground cover variables were also sampled to determine how variation in ant populations was correlated with differences in vegetation and ground cover. Ant species richness remained unchanged for all treatments. Tree cover had no significant effect on ant populations. Significantly more ants were trapped after the burn treatment on burn plots. Variation in ant populations was not directly correlated with any of the vegetation or ground cover variables. According to ANOVA and multivariate analyses, elevation had the greatest effect on changes in ant communities, likely due to increased moisture availability. Our results suggest that management for conservation of sagebrush-steppe ecosystems in this and similar watersheds should include a range of elevations to ensure maximum ant species diversity.     

Winter and early spring bird communites in grasslands, shrubsteppe, and juniper woodlands in central Oregon

We compared winter (December, January, and February) and early spring (March and April) bird communities among 4 successional stages that included grassland, shrubsteppe, juniper-shrubsteppe, and old-growth juniper woodland in central Oregon. Birds were surveyed monthly from December through April in 1998/1999 and 1999/2000 using the point count method to estimate relative abundance of birds (individuals ? transect –1 year –1 ). We used distance sampling to correct for potential bias in estimating abundance and density among successional stages. A total of 4513 birds (32 species) were detected. Relative abundance of total birds (all species combined) was similar in the juniper-shrubsteppe and old-growth woodland (48.7 and 48.9 individuals ? transect –1 , respectively) but 6 times greater than in the grassland and shrubsteppe (8.0 and 7.7 individuals ? transect –1 , respectively) during the 5-month period. Total density of birds was greater in old-growth juniper woodland and juniper-shrubsteppe than in grassland or shrubsteppe. However, median bird species richness in 1998/1999 was highest in grassland and lowest both in shrubsteppe and juniper-shrubsteppe, and in 1999/2000 it was highest in shrubsteppe and lowest in juniper-shrubsteppe and oldgrowth juniper woodland. American Robins and Townsend's Solitaires were the most abundant species in junipershrubsteppe and old-growth juniper woodland during the winter months. Sage Sparrows and Horned Larks were the most abundant species in shrubsteppe during winter, and Horned Larks were most abundant in grasslands during the early spring transition period prior to nesting. Our results indicate that a different suite of species use these successional stages during the nonbreeding season. If avifauna conservation is a part of long-term management goals, a broad range of successional stages should be maintained on the landscape to provide habitat for a variety of avian species throughout the year.     

Birds of northern Black Mesa, Navajo County, Arizona

Two hundred forty-one species of birds have been identified from northern Black Mesa, Arizona. This region's avifauna was poorly known until the late 1970s when large-scale coal mining began. Vegetation of the region is predominantly Great Basin desert scrub, pinyon-juniper woodland, and mixed-conifer woodland. The latter vegetation supports an assemblage of isolated montane bird species unique to the region. Numerous environmental changes have recently affected the bird life of Black Mesa. These include large-scale type conversions (pinyon-juniper clearing and surface mining), pond construction, and establishment of exotic vegetation.     

Brood parasitism and nest survival of Brown-headed Cowbird hosts at high-elevation riparian sites in the eastern Sierra Nevada, California

Deciduous riparian ecosystems in the western United States provide habitat for a higher density of breeding birds than reported for any other avian habitat type and provide habitat for more breeding bird species than adjacent uplands. On the east slope of the Sierra Nevada, riparian ecosystems make up Molothrus ater )—an obligate brood parasite that forages on bare ground and feedlots but typically commutes to distinct shrubland or woodland habitats for breeding. We examined nest survival, brood parasitism, breeding phenology, and causes of nest failure for birds at North Lake and Rock Creek: 2 high-elevation (>2500 m) riparian breeding habitats adjacent to recreational development and within cowbird commuting distance to additional potential foraging sites. Nest survival tended to be higher for host species at Rock Creek than for those at North Lake, but parasitism rates were not significantly different between plots. Of 21 open-cup nesting species, 12 were parasitized. We found the highest rate of parasitism (92%) for Warbling Vireos ( Vireo gilvus ) at North Lake, and parasitism contributed to lower total nest survival there (14%). For nearly all species, parasitized nests were less successful and produced fewer young than nonparasitized nests. However, predation was the leading cause of complete nest failure across all species and contributed to the lowest total nest survival estimates for Western Wood-Pewees ( Contopus sordidulus, 11%) and Dusky Flycatchers ( Empidonax oberholseri, 15%) at North Lake and for Dark-eyed Juncos ( Junco hyemalis, 15%) at Rock Creek. Nest survival was relatively high for Western Wood-Pewees (41%) at Rock Creek and for Yellow Warblers ( Dendroica petechia, 47%) at North Lake. We noted whether the arrival of pack animals at pack-station corrals contributed to variation in cowbird numbers at corrals or in parasitism rates at the 2 sites. Cowbirds occupied corrals before and after pack-stock arrival, and most host clutches were completed prior to pack-stock arrival at nearby corrals, suggesting that the presence of pack animals did not directly affect cowbird host species.     

Habitat use by small mammals in southeastern Utah, with reference to Mexican Spotted Owl management

We determined temporal and spatial differences in abundance and habitat use by small mammals in southeastern Utah as part of an effort to enhance management of the Mexican Spotted Owl ( Strix occidentalis lucida ), listed by the federal government as threatened. Woodrats ( Neotoma spp.) were captured only in canyons and most frequently in the pinyon-juniper ( Pinus edulis-Juniperus osteosperma ) vegetation type. White-footed mice ( Peromyscus spp.) were found in a variety of vegetation types in both canyons and mesas. The deer mouse ( P. maniculatus ) was generally the most frequently captured species among vegetation types. We found seasonal and yearly differences in relative abundance of each small mammal species. Our data suggest that the pinyon-juniper vegetation type within canyons is an important component of Mexican Spotted Owl habitat.     

Riparian vegetation along the middle Snake River, Idaho: zonation, geographical trends, and historical changes

A baseline study was conducted on an 83-km free-flowing reach of the Snake River between Swan Falls Dam and the Idaho-Oregon border. The research had 2 components: (1) field characterization and inventory of existing riparian flora, vegetation, and environment (soils, topography, streamflow), and (2) determination and mapping, using a geographic information system, of historic changes in riparian vegetation based on a time series (1938-39, 1957, 1969, 1987) of aerial photographs. The flora was diverse, with 185 species of vascular plants identified, 63 of which were exotics. Vegetation was structured vertically along the riverbank gradient into lifeform-defined habitat types: emergent, riparian shrub-forb, tree, transitional grass-shrub, and upland. Riverbank seepage, probably of agricultural origin, blurred zonation patterns on some sites and added species to the overall flora. Upstream-downstream differences existed in the physical characteristics and vegetation of river subreaches. Coverage of riparian woodland, island riparian and total riparian vegetation, and area of islands increased since the 1930s, with the greatest changes in the 1969-1987 interval. Possible contributing facts were (1) significant declines in annual minimum flows since the 1950s, (2) decreases in peak flows following the completion of Swan Falls Dam and some upstream dams since the 1920s, (3) introduction and spread of exotic tree species ( Elaeagnus angustifolia and Tamarix spp.) and (4) possible effects of intensive agriculture on river sediment load and soil nutrients. The introduction and proliferation of purple loosestrife ( Lythrum salicaria ) could have considerable future effects on vegetation-channel dynamics in the middle Snake River.     

Comparative habitat use in a juniper woodland bird community

We compared vegetation structure used by 14 bird species during the 1998 and 1999 breeding seasons to determine what habitat features best accounted for habitat division and community organization in Utah juniper ( Juniperus osteosperma ) woodlands of southwestern Wyoming. Habitat use was quantified by measuring 24 habitat variables in 461 bird-centered quadrats, each 0.04 ha in size. Using discriminant function analysis, we differentiated between habitat used by 14 bird species along 3 habitat dimensions: (1) variation in shrub cover, overstory juniper cover, mature tree density, understory height, and decadent tree density; (2) a gradient composed of elevation and forb cover; and (3) variation in grass cover, tree height, seedling/sapling cover, and bare ground/rock cover. Of 14 species considered, 9 exhibited substantial habitat partitioning: Mourning Dove ( Zenaida macroura ), Bewick's Wren ( Thryomanes bewickii ), Blue-gray Gnatcatcher ( Polioptila caerulea ), Mountain Bluebird ( Sialia currucoides ), Plumbeous Vireo ( Vireo plumbeus ), Green-tailed Towhee ( Pipilo chlorurus ), Brewer's Sparrow ( Spizella breweri ), Dark-eyed Junco ( Junco hyemalis ), and Cassin's Finch ( Carpodacus cassinii ). Our results indicate juniper bird communities of southwestern Wyoming are organized along a 3-dimensional habitat gradient composed of woodland maturity, elevation, and juniper recruitment. Because juniper birds partition habitat along successional and altitudinal gradients, indiscriminate woodland clearing as well as continued fire suppression will alter species composition. Restoration efforts should ensure that all successional stages of juniper woodland are present on the landscape.     

Red Butte Canyon Research Natural Area: history, flora, geology, climate, and ecology

Red Butte Canyon is a protected, near pristine canyon entering Salt Lake Valley, Utah. It contains a well-developed riparian zone and a perennial stream; hillside vegetation ranges from grasslands on the lower limits to Douglas-fir and aspen stands at the upper elevations. In this paper we describe the history of human impact, natural history aspects of climate, geology, and ecology, and faunal and floral information for key species in the canyon. The role and importance of Research Natural Areas is discussed, particularly with respect to the need to protect Red Butte Canyon:one of the few remaining undisturbed riparian ecosystems in the Intermountain West.     

Bait,habitat preferences and the phenology of some lesser dung flies (Diptera: Sphaeroceridae) in Britain

Two thousand eight hundred and sixty Sphaeroceridae representing 45 species were caught in eight traps, baited with horse, cow or hamster dung or carrion, placed in woodland and in the open at Silwood Park, Berkshire, during 1983/4. A list of the species and the numbers of each caught in each of the traps is given. Comparisons between the catches of open and woodland habitats and between the catches of the different baits used are made. The habitat and bait preferences and the phenologies of the dominant species are discussed.     

Bird use of riparian vegetation along the Truckee River, Califonia and Nevada

The Truckee River in California and Nevada is subject to diverse water regimes and a corresponding variety of flow rates. Original riparian vegetation has been altered by these variable flow rates and by a variety of human uses resulting in loss of native riparian vegetation from its historic extent. We conducted bird surveys along the Truckee River during spring 193 to (1) determine relationships between birds and the present vegetation; (2) determine the importance of different vegetation types to sensitive bird species that have declined recently in the western United States due to competition from exotic plant species, cowbird ( Molothrus ater ) parasitism, reduction in nesting habitat, or other unidentified reasons; and (3) establish a monitoring program and collect baseline data for future comparisons. The most frequently detected bird species throughout the study was the Brown-headed Cowbird. The greatest number of bird species (98 of 116) was found in the native mixed willow ( Salix spp.) riparian scrub vegetation type. We recommend protecting the remaining native riparian vegetation types for bird habitat along the Truckee River.     

Evaluating plant invasions from both habitat and species perspectives

We present an approach to quantitatively assess nonnative plant invasions at landscape scales from both habitat and species perspectives. Our case study included 34 nonnative species found in 142 plots (0.1 ha) in 14 vegetation types within the Grand Staircase-Escalante National Monument, Utah. A plot invasion index, based on nonnative species richness and cover, showed that only 16 of 142 plots were heavily invaded. A species invasive index, based on frequency, cover, and number of vegetation types invaded, showed that only 7 of 34 plant species were highly invasive. Multiple regressions using habitat characteristics (moisture index, elevation, soil P, native species richness, maximum crust development class, bare ground, and rock) explained 60% of variation in nonnative species richness and 46% of variation in nonnative species cover. Three mesic habitats (aspen, wet meadow, and perennial riparian types) were particularly invaded (31 of 34 nonnative species studied were found in these types). Species-specific logistic regression models for the 7 most invasive species correctly predicted occurrence 89% of the time on average (from 80% for Bromus tectorum , a habitat generalist, to 93% for Tamarix spp., a habitat specialist). Even with such a modest sampling intensity (<0.1% of the landscape), this multiscale sampling scheme was effective at evaluating habitat vulnerability to invasion and the occurrence of the 7 most invasive nonnative species. This approach could be applied in other natural areas to develop strategies to document invasive species and invaded habitats.     

Terrestrial vertabrates of Scotts Bluff National Monument, Nebraska

A survey of terrestrial vertebrates was conducted at the Scotts Bluff National Monument (SBNM), Nebraska, to determine species composition, relative abundance, and distribution by habitat of the existing amphibians, reptiles, birds, and mammals. Various sampling methods were used to detect animal species richness and to estimate population densities. The seven major habitat types at SBNM contained 4 species of amphibians, 8 reptiles, 96 birds, and 28 mammals. The only endangered or threatened species observed was a dead bald eagle ( Haliaeetus leucocephalus ). The racer ( Coluber constrictor ) and prairie rattlesnake ( Crotalus viridis ) were the two most abundant snakes. The prairie falcon ( Falco mexicanus ) was a common raptor that nested in Scotts Bluff. Sympatric populations of mule deer ( Odocoileus hemionus ) and white-tailed deer ( Odocoileus virginianus ) appeared to be hybridizing. The riverine woodland contained the greatest number of fauna species with only 4% surface area of SBNM. Sharp-tailed grouse ( Tympanuchus phasianellus ) and pronghorn antelope ( Antilocapra americana ), absent from the Monument, are two potential native species for reintroduction.     

Distribution and abundance of birds in the White Mountains, California

This paper quantifies the distribution and abundance of birds in the White Mountains, Inyo and Mono counties, California, during spring-summer 1989-91, to establish a baseline for monitoring the area's avifauna. Overall 58 species were encountered in the single-leaf pinyon-Utah juniper ( Pinus monophylla-Juniperus osteosperma ) zone, and 61 species in the bristlecone-limber pine ( P. longaeva-P. flexilis ) zone. The bristlecone-limber pine zone had a significantly greater overall bird abundance relative to the pinyon-juniper. Both zones were characterized by few very abundant species, a few moderately abundant species, and numerous rare species. The Black-throated Gray Warbler (scientific names in tables), Gray Flycatcher, and Pinyon Jay were the most abundant species in the pinyon-juniper whereas the Clark's Nutcracker, Mountain Chickadee, and Cassin's Finch were the most abundant species in the bristlecone-limber pine. There were few ecological or taxonomic replacements of species between zones, with the differences in distribution and abundance related primarily to the interaction between elevation and vegetation. Significant inter-year variation in abundance was found for about 20 species in each zone—more species showed declining rather than increasing trends. The Mountain Chickadee and White-breasted Nuthatch declined, whereas the Gray Flycatcher and Rock Wren increased across years in both zones. Reasons for declines in some species might be the severe drought that continued throughout this study.     

Plant community changes within a mature pinyon-juniper woodland

Vegetal composition was determined during 1974 and 1984 using 60 permanent 50 m 2 plots within a mature pinyon-juniper community in northeastern Utah. Results indicated that not only was there little significant change in community composition, but with many species frequency and density remained nearly the same during the decade.     

Fat depth at the xiphoid process--a rapid index to deer condition

Vegetal composition was determined during 1974 and 1984 using 60 permanent 50 m 2 plots within a mature pinyon-juniper community in northeastern Utah. Results indicated that not only was there little significant change in community composition, but with many species frequency and density remained nearly the same during the decade.     

Habitat characteristics of small mammals in southeastern Utah

We describe habitat characteristics of 8 small mammal species occurring in the Manti-LaSal National Forest in southeastern Utah. Thirty-seven grids across 7 vegetation types were sample by live-trapping from May through October 1994 and 1995. Logistic regression models of habitat measurements correctly classified small mammal presence 36%-87% of the time. Except for 1 instance, 4 Peromyscus species present in a specific vegetation type were negatively associated with forb or grass cover. Microtus montanus was present primarily on mesas and was positively associated with tall tree and low shrub cover. Neotoma mexicana was present only in canyons and was found primarily in the pinyon-juniper ( Pinus edulis-Juniperus osteosperma ) vegetation type. Tamias spp. was present in all vegetation types. Our results allow better management of these species by providing greater understanding of their use of habitat within vegetation types.