Revision of the smiling worms,genera Prosadenoporus Bürger, 1890 and Pantinonemertes Moore and Gibson, 1981 and description of a new species Prosadenoporus floridensis sp. nov. (Prosorhochmidae; Hoplonemertea; Nemertea) from Florida and Belize

The hoplonemertean genera Prosadenoporus Bürger, 1890 Bürger, O. 1890. Untersuchungen über die anatomie und histologie der nemertinen nebst beiträgen zur systematik [Studies of anatomy and histology of nemerteans with contribution to systematics].. Z Wiss Zool., 50: 1–277. German [Google Scholar] and Pantinonemertes Moore and Gibson, 1981 Moore, J. and Gibson, R. 1981. The Geonemertes problem (Nemertea).. J Zool., 194: 175–201.  [Google Scholar] are revised and synonymized based on a morphological re‐evaluation. We redefine Prosadenoporus Bürger, 1890 Bürger, O. 1890. Untersuchungen über die anatomie und histologie der nemertinen nebst beiträgen zur systematik [Studies of anatomy and histology of nemerteans with contribution to systematics].. Z Wiss Zool., 50: 1–277. German [Google Scholar] on the basis of characters held in common by the eight species: Prosadenoporus agricola (Willemoes‐Suhm, 1874 Willemoes‐Suhm, R. 1874. On a land‐nemertean found in the Bermudas.. Ann Mag Nat Hist. Ser 4., 13: 409–411.  [Google Scholar]) comb. nov., Prosadenoporus arenarius Bürger, 1890 Bürger, O. 1890. Untersuchungen über die anatomie und histologie der nemertinen nebst beiträgen zur systematik [Studies of anatomy and histology of nemerteans with contribution to systematics].. Z Wiss Zool., 50: 1–277. German [Google Scholar], Prosadenoporus spectaculum (Yamaoka, 1940 Yamaoka, T. 1940. Two nemerteans from the Riukiu Islands.. Annot Zool Japon., 19: 13–18.  [Google Scholar]) comb. nov., Prosadenoporus winsori (Moore and Gibson, 1981 Moore, J. and Gibson, R. 1981. The Geonemertes problem (Nemertea).. J Zool., 194: 175–201.  [Google Scholar]) comb. nov., Prosadenoporus enalios (Moore and Gibson, 1981 Moore, J. and Gibson, R. 1981. The Geonemertes problem (Nemertea).. J Zool., 194: 175–201.  [Google Scholar]) comb. nov., Prosadenoporus mooreae (Gibson, 1982b Gibson, R. 1982b. Nemerteans of the Great Barrier Reef. 5. Enopla Hoplonemertea (Monostilifera).. Zool J Linn Soc., 75: 269–296.  [Google Scholar]) comb. nov., Prosadenoporus mortoni (Gibson, 1990 Gibson, R. 1990. “The macrobenthic nemertean fauna of Hong Kong.”. In The Marine Flora and Fauna of Hong Kong and Southern China, Edited by: Morton, B. 33–212. Hong Kong: Hong Kong University Press.  [Google Scholar]) comb. nov. and Prosadenoporus fujianensis (Sun, 2001 Sun, S. 2001. A new mangrove‐dwelling nemertean from China.. Hydrobiologia, 456: 199–209.  [Google Scholar]) comb. nov. We describe a new semi‐terrestrial species Prosadenoporus floridensis sp. nov. from Belize and Florida and compare its morphology to other species of Prosadenoporus. The average sequence divergence of P. floridensis sp. nov. from other congeners is 9.15% (16S) and 10.65% (COI) and 7.8% and 10.3% respectively from the nearest sequenced congener P. mortoni.     

Revision of Flabelliderma Hartman, 1969 (Polychaeta: Flabelligeridae)

The flabelligerid genus Flabelliderma Hartman, 1969 Hartman, O. 1969. Atlas of the sedentariate polychaetous Annelids from California, 812Los Angeles: University of Southern California, Allan Hancock Foundation.  [Google Scholar] is redefined and its type species is re‐established. In comparison with Flabelligera Sars, 1829 Sars, M. 1829. Bidrag til Söedyrenes Naturhistorie; Förste Hæfte, 60Bergen: Dahl.  [Google Scholar], the diagnostic features include the fusion of papillae forming tubercles, often adhering sediment particles, and notopodial lobes forming large ovoid lobes. Four species are described and presented as new combinations, and three others were previously undescribed; thus the species included are F. papillosa (Essenberg, 1922 Essenberg, C. E. 1922. Stylarioides papillosa, sp. nov., a new annelid from the San Diego region.. University of California, Publications in Zoology, 22: 379–381.  [Google Scholar]) from southern California, F. berkeleyorum n. sp. from Washington, F. claparedei (de Saint‐Joseph, 1898 de Saint‐Joseph, A. 1898. Les annélides polychètes des côtes de France (Manche et Océan).. Annales des Sciences Naturelles, Zoologie, Série 8, 5: 209–464.  [Google Scholar]) from the Bay of Biscay, F. gourdoni (Gravier, 1906 Gravier, C. 1906. Sur les annélides polychètes recueillies par l'Expédition Antarctique Française (Aphroditiens, Amphinomiens, Flabelligériens, Maldaniens, Ampharétiens).. Bulletin du Museum d'Histoire Naturelle, Paris, 7: 535–540.  [Google Scholar]) from Antarctica, F. lighti n. sp. from Guadalupe Island, western Mexico, F. ockeri n. sp. from southern California, and F. pruvoti (Fauvel, 1930 Fauvel, P. 1927. Polychètes sédentaires & addenda aux polychètes errantes.. Faune de France, 16: 1–494.  [Google Scholar]) from New Caledonia.     

Revision of the hirsuticornis‐like species of Macrothrix Baird, 1843 (Cladocera: Anomopoda: Macrothricidae) from Subantarctic and temperate regions of the southern hemisphere

The aim of this paper is to revise populations of Macrothrix cf. hirsuticornis (Cladocera: Anomopoda: Macrothricidae) from different regions of the southern hemisphere. It is demonstrated that M. hirsuticornis Norman and Brady, 1867 Norman, A. M. and Brady, G. S. 1867. A monograph of the British Entomostraca belonging to the families Bosminidae, Macrothricidae and Lynceidae.. Natural History Transactions of Northumberland and Durham, 1: 354–408.  [Google Scholar] s. str. is absent there, and five related species occupy different Subantarctic islands and the southernmost portions of South America, and Africa. Macrothrix boergeni Studer, 1878 Studer, T. 1878. Beiträge zur Naturgeschichte wirbelloser Tiere von Kerguelensland.. Archiv für Naturgeschichte, 44: 102–121.  [Google Scholar] from the Kerguelen Archipelago is redescribed and a neotype is selected. All populations in the southernmost portion of continental South America, Tierra del Fuego, Falklands, South Georgia, South Orkney Islands, and on the Antarctic Peninsula belong to M. oviformis Ekman, 1900 Ekman, S. 1900. Cladoceren aus Patagonien, gesammelt von der schwedischen Expedition nach Patagonien 1899.. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 14: 62–84.  [Google Scholar]. All the taxa described from this region—M. ciliata Vávra, 1900 Vávra, W. 1900. “Süsswasser‐Cladoceren. Ergebnisse der Hamburger magalhaensische Sammelreise 1892/93. II Band. Arthropoden.”. In Hamburg 1–25.  [Google Scholar], M. odontocephala Daday, 1902 Daday, E. 1902. Mikroskopische Süsswasserthiere aus Patagonien, gesammelt von Dr. Filippo Sylvestri.. Természetrajzi Füzetek, Budapest, 25: 201–310.  [Google Scholar], M. propinqua Sars, 1909 Sars, G. O. 1909. Fresh‐water Entomostraca from South Georgia.. Archiv for Mathematik og Naturvidenskab, 30: 1–35.  [Google Scholar], and, probably, M. inflata Daday, 1902 Daday, E. 1902. Mikroskopische Süsswasserthiere aus Patagonien, gesammelt von Dr. Filippo Sylvestri.. Természetrajzi Füzetek, Budapest, 25: 201–310.  [Google Scholar]—are junior synonyms of M. oviformis. Two new species are established: M. sarsi sp. nov. from the Cape region of South Africa and M. ruehei sp. nov. from Crozet, Marion islands, and Île Amsterdam. Macrothrix cf. flagellata Smirnov and Timms, 1983 Smirnov, N. N. and Timms, B. V. 1983. A revision of the Australian Cladocera (Crustacea).. Records of the Australian Museum, Supplement, 1: 1–132. [Crossref] , [Google Scholar], previously known only from Tasmania, is found on Macquarie Island too. Differences between species from the southern hemisphere and Palaearctic M. hirsuticornis are summarized. It is demonstrated that characters of the general body shape (i.e. presence of a hood or a tooth on posterior head border) have a limited value for the systematics of Macrothrix. In contrast, some fine details, mostly missed by previous authors, are valuable for species discrimination. The present study increases the number of species recorded from the Antarctic‐Subantarctic region. Probably, the current pattern of Macrothrix distribution results from a disruption of a pan‐continental (early Mesozoic?) species complex.     

The diverse Grania fauna (Clitellata: Enchytraeidae) of the Esperance area,Western Australia,with descriptions of two new species

Seven species of the marine enchytraeid genus Grania Southern, 1913 are described from sediments sampled during the 2003 International Workshop on the Marine Flora and Fauna of Esperance Bay and the Recherche Archipelago, on the southern coast of Western Australia. Two species are new to science, the euryhaline Tasmanian G. dolichura Rota and Erséus, 2000 Rota, E. and Erséus, C. 2000. Two new and peculiar species of Grania (Annelida: Clitellata: Enchytraeidae) inhabiting Tasmanian estuaries.. New Zealand Journal of Zoology, 27: 245–254. [Taylor & Francis Online], [Web of Science ®] , [Google Scholar] represents a new record for the state, and the remaining four species were known from other parts of Western Australia. Grania quaerens sp. n. is recognized by having a high chaetal index ( = short chaetal foot), small coelomocytes, penial apparati with long whip‐like terminal stylets, conspicuous spermathecae with ectally bulbous ducts, and ectally granulated ampullae housing sperm rings in their ental region. Grania sperantia sp. n. is readily distinguishable by the complete lack of lateral chaetae, a multiple‐banded pattern of the clitellum, extremely long sperm funnels, and the intrasegmental location of the spermathecal pores. The latter new species and four others in the collection (G. bykane Coates, 1990 Coates, K. A. 1990. “Marine Enchytraeidae (Oligochaeta, Annelida) of the Albany area, Western Australia.”. In The marine flora and fauna of Albany, Western Australia, 1, Edited by: Wells, F. E, Walker, D. I, Kirkman, H and Lethbridge, R. 13–41. Perth: Western Australian Museum.  [Google Scholar], G. crassiducta Coates, 1990 Coates, K. A. 1990. “Marine Enchytraeidae (Oligochaeta, Annelida) of the Albany area, Western Australia.”. In The marine flora and fauna of Albany, Western Australia, 1, Edited by: Wells, F. E, Walker, D. I, Kirkman, H and Lethbridge, R. 13–41. Perth: Western Australian Museum.  [Google Scholar], G. dolichura, and G. ersei Coates, 1990 Coates, K. A. 1990. “Marine Enchytraeidae (Oligochaeta, Annelida) of the Albany area, Western Australia.”. In The marine flora and fauna of Albany, Western Australia, 1, Edited by: Wells, F. E, Walker, D. I, Kirkman, H and Lethbridge, R. 13–41. Perth: Western Australian Museum.  [Google Scholar]) are remarkable in possessing the head organ, a sensory structure unique to Grania that was not noted previously in Western Australian species. When considering the whole genus, the geographic pattern of the head organ appears southern‐centred: of the 17 species of Grania reported to possess it, as many as 13 inhabit the southern latitudes. The seventh species of the Esperance collection, G. vacivasa Coates and Stacey, 1993 Coates, K. A. and Stacey, D. 1993. “The marine Enchytraeidae (Oligochaeta, Annelida) of Rottnest Island, Western Australia.”. In The marine flora and fauna of Rottnest Island, Western Australia, Edited by: Wells, F. E, Walker, D. I, Kirkman, H and Lethbridge, R. 391–414. Perth: Western Australian Museum.  [Google Scholar], is notable for the kind of items found in its gut and the unusual appearance of its pygidium.     

Taxonomy of the genus Gabrius Stephens, 1829 (Coleoptera: Staphylinidae: Philonthina) from China,with description of two new species

In this paper, two new species of the genus Gabrius from China are described and illustrated: G. zhoui sp. nov. from Sichuan and G. invisus sp. nov. from Beijing, Hubei, Hunan, Henan, Shaanxi, Sichuan and Zhejiang. Gabrius imitator (Fauvel, 1895 Fauvel, A. 1895. Staphylinides nouveaux de l'Inde et de la Malaisie. Rev Entomol, 14: 180–286.  [Google Scholar]), G. lebedevi (Bernhauer, 1910 Bernhauer, M. 1910. Zur Staphylinidenfauna des palaearktischen Gebietes. Soc Entomol, 25: 71–71. 78–79 [Google Scholar]), G. tirolensis (Luze, 1903 Luze, G. 1903. Eine neue Art der Staphyliniden-Gattung Philonthus Curtis aus Mitteleuropa. Verhandlungen der k. k. Zool-botan Gesellschaft Wien, 53: 386 [Google Scholar]), G. kambaitiensis Scheerpeltz, 1965 Scheerpeltz, O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except Megalopsidiinae et Steninae). Ark Zool, 17: 93–371.  [Google Scholar], G. cinctiventris Scheerpeltz, 1965 Scheerpeltz, O. 1965. Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except Megalopsidiinae et Steninae). Ark Zool, 17: 93–371.  [Google Scholar], G. egens (Sharp, 1874 Sharp, DS. 1874. The Staphylinidae of Japan. Entomol Soc London Trans, : 1–103.  [Google Scholar]), G. smetanai Schillhammer, 1997 Schillhammer, H. 1997. Taxonomic revision of the Oriental species of Gabrius Stephens (Coleoptera: Staphylinidae). Mono Coleop, 1: 1–139.  [Google Scholar] and G. zerchei Schillhammer, 1996 Schillhammer, H. 1996. New genera and species of Asian Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Koleopterol Rundsch, 66: 59–71.  [Google Scholar] are reported from China for the first time. Thus, a total of 53 species of the genus Gabrius are known from China.     

Notes on four corinnid species from Korea,with the description of Trachelas joopili new species (Arachnida: Araneae: Corinnidae)

Four species and two genera of corinnid spiders are recognized from Korea and are described and illustrated. Castianeira shaxianensis Gong, 1983 Gong, J. X. 1983. Neue und wenig bekannte Clubionidae aus China (Arachnida: Araneae).. Verh Natur Vereins Hamburg., 26: 61–68.  [Google Scholar] has previously been synonymized with C. paikdoensis Kim, 1997 Kim, J. P. 1997. A new species of the genus Castianeira (Araneae: Corinnidae) from Korea.. Korean Arachnol., 13(1): 1–5.  [Google Scholar] and is misidentified with C. flavimaculata Hu, Song and Zheng, 1985 in Korea. Trachelas acuminus (Zhu and An, 1988 Zhu, M. S. and An, R. 1988. Two new species of the genus Clubiona from China (Araneae: Clubionidae).. J Hebei Norm Univ (Nat. Sci. Ed.)., 1988: 72–75.  [Google Scholar]) is synonymized with Trachelas coreanus Paik, 1991 based on the copualtory duct, the position of the spermatheca and the shape of the circular genital opening of the female situated posteriorly near the epigastric furrow. Trachelas joopili, a new species, is described and is distinguished by the largely triangular apophysis and slender coiled modified cymbium at the basal and distal part, a large horn‐shaped tibial apophysis, and slender linear embolus, twice coiled along with slender distal cymbium. A map showing the distribution of these species and a key to the species of two genera (Castianeira and Trachelas) from Korea are provided.     

The millipede tribe Nedyopodini,with special reference to the fauna of Taiwan (Diplopoda: Polydesmida: Paradoxosomatidae)

The east and southeast Asian millipede tribe Nedyopodini is here restricted to a single genus, Nedyopus Attems, 1914 Attems, C. G. 1914. Die indo‐australischen Myriopoden.. Archiv für Naturgeschichte, 80: 1–398.  [Google Scholar]; the following genera are here considered to be junior subjective synonyms: Vaulogerodesmus Brölemann, 1916 Brölemann, H. W. 1916. Essai de classification des polydesmiens (Myriapodes).. Annales de la Société Entomologique de France, 84: 523–607.  [Google Scholar], Nesodyopus Verhoeff, 1940 Verhoeff, K. W. 1940. Zur Kenntnis ostasiatischer Diplopoden V (Polydesmoidea).. Zoologischer Anzeiger, 131: 129–145.  [Google Scholar], n. syn., and Varyomorpha Wang, 1957, n. syn. This taxonomically confused tribe/genus is reviewed and redefined. Based on new collections, the fauna of Taiwan currently contains five Nedyopus species including the redescribed N. hsientienensis (Wang, 1957) and N. pectinatus (Wang, 1957), both n. comb. ex Varyomorpha, as well as the newly described N. caudatus n. sp., N. latus n. sp., and N. wui n. sp. A key to these five species is provided.