The first twisted-wing parasitoid in Eocene amber from north-eastern China (Strepsiptera: Myrmecolacidae)

An Early Eocene amber twisted-wing parasitoid (Strepsiptera) is described and figured from the Fushun Coalfield in north-eastern China, representing the first record of this order for the Tertiary of Asia and the second extinct genus of Myrmecolacidae. Kronomyrmecolax fushunicus gen. et sp. nov. is represented by a single male which is distinguished from other living and fossil myrmecolacids, particularly Palaeomyrmecolax Kulicka, in middle Eocene Baltic amber. The genera differ in details of hind wing venation and antennal morphology.     

First described fossil representatives of the parasitoid wasp taxa Asaphesinae n. n. and Eunotinae (Hymenoptera: Chalcidoidea: Pteromalidae sensu lato) from Eocene Baltic amber

ABSTRACT

New Baltic amber species of Pteromalidae sensu lato are described, from two different subfamilies, Asaphesinae n. n. and Eunotinae. Asaphesinae is provided as a replacement name for Asaphinae Ashmead 1904, which is a junior homonym of the trilobite family Asaphidae Burmeister 1843. Coriotela lasallei gen. n., sp. n.. and Butiokeras costae gen. n., sp. n.. are described as the first known fossil species of Asaphesinae and Eunotinae, respectively. These species establish the minimum known age of both groups in the Eocene. Taxonomic changes are also proposed for some extant species. The genus Desantisiana Neder de Román syn. n.. is found to be a junior synonym of Notoglyptus, and its only described species is transferred as Notoglyptus jujuyensis (Neder de Román) comb. n.. The tribe Calyconotiscini, previously classified in Eunotinae, is abolished and Calyconotiscus Narendran & Saleem is transferred to Pireninae.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:7A107FF9-28E7-40AA-8A9B-71321E476C07     

A new genus and species of Ricaniidae from Palaeocene deposits in North Dakota

Summary

A new genus and species, Cotradechites lithinus Fennah, are proposed for a fossil tegmen of a Ricaniid (Homoptera : Fulgoroidea) collected in clay deposits of Latest Palaeocene age near Dickinson, North Dakota. The new genus is considered to be most nearly related to the modern tropical American genera Cotrades and Semestra.     

A redescription of the fossil coleoid cephalopod genus Palaeololigo Naef, 1921 (Decapodiformes: Palaeololiginidae) and its relationship to Recent squids

Neocoleoid cephalopods are poorly represented in the fossil record and fossils which can be confidently aligned with extant species are very scarce. The rare Solnhofen genus Palaeololigo is an exception to this, showing marked resemblance to Recent squids. Naef misinterpreted the structure of the fossilized gladius of Palaeololigo and the present study redescribes the genus. The gladius and fossilized fin impressions of Palaeololigo closely resemble that of the extant decapodiform squid genera Bathyteuthis and Chtenopteryx (Superfamily Bathyteuthoidea). The possible relationship between Palaeololigo and the bathyteuthoids and their position in decapodiform phylogenetics is discussed.     

A new genus of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to the Brazilian Atlantic Forest

ABSTRACT

A new genus of the tribe Deltochilini, Atlantemolanum new genus, endemic to the Atlantic Forest of Brazil, is proposed here to include two species formerly described in the genus Deltochilum based on the unique morphologic characters shared by Deltochilum riehli = Atlantemolanum riehli (Harold 1868) new combination and Deltochilum costalimai = Atlantemolanum costalimai (Pereira and D’Andretta 1955) new combination.

http://www.zoobank.org/urn:lsid:zoobank.org:act:7BBD89C3-2F23-4CEF-9560-C29695631E04     

A re-assessment of the genera Holorusia Loew (=Ctenacroscelis Enderlein), Ischnotoma Skuse and Zelandotipula Alexander (Diptera: Tipulidae), with notes on their phylogeny and zoogeography

Summary

A short historical background to the genera considered is given. A review is made of a number of characters which had previously only been treated inconsistently, or not at all. As a result, the old-world genus Ctenacroscelis is merged with Holorusia sensu stricto, as represented by the isolated Californian type-species. All other species previously referred to Holorusia sensu lato are redistributed to two genera; the tropical South American and New Zealand forms are united into Zelandotipula; the Chilean element and the Australian species are united into Ischnotoma, the name Icriomastax Enderlein being resurrected as a subgenus to receive most of the Chilean species.

Tentative phylogenetic and zoogeographic observations are made. Holorusia and Ischnotoma are considered and discussed as sister-groups, whilst Zelandotipula is believed to be only distantly related. Zoogeographically Holorusia is seen as a successful Asiatic genus; Ischnotoma as a receding group, with extreme southern-continental distribution, having had a much wider distribution as indicated by the European Oligocene fossil Holorusia vasifera, transferred here to Ischnotoma. Zelandotipula is viewed as an essentially Neotropical genus, which has made one ‘invasion’ of New Zealand, probably in the geologically recent past.

Résumés of the principal features of the re-defined genera are given, together with systematic notes on certain species, and species catalogues giving distributions and all new name combinations.     

Revision of the Palaearctic species of the genus Clinocentrus (Hymenoptera,Braconidae)

Revision of the Palaearctic species of the genus Clinocentrus Haliday is given. The composition and geographic distribution of the genus Clinocentrus and the probable generic position of fossil species described by Brues (1933) as Clinocentrus are discussed. The systematic position of this genus in Exothecinae is also discussed and arguments presented to support the differentiation of exothecines and rogadines. Nine species of Clinocentrus are figured and re-described. Three new species and two new subspecies are figured and described: C. kalmyk from Kalmykia, C. kozlovi from Mongolia, C. rhysipoloides from Primorsk Territory of Russia, C. umbratilis ssp. disruptus from Taiwan and C. vestigator ssp. orientalis from Primorsk Territory of Russia. C. stigmaticus Marshall, 1897 and C. jaroshevskyi Telenga, 1941 are synonymized with C. vestigator (Haliday, 1836); C. tarsalis Ashmead, 1894—with C. exsertor (Nees, 1812) and C. gracilipes (Thomson, 1891)—with C. cunctator (Haliday, 1836), Lectotypes of Exothecus analis Wesmael, 1838 and C. antefurcalis Granger, 1949 are designated. The new combination is given: Rhysipolis antefurcalis (Granger). A key to 12 Palaearctic species of Clinocentrus is given.     

An Early Cretaceous anomopod (Crustacea: Branchiopoda) preserved in amber that reveals an unexpected venture during the evolution of the order

An anomopod branchiopod crustacean that displays many structural similarities to the extant genera Scapholeberis and Megafenestra (family Daphniidae), fortuitously preserved in amber of Early Cretaceous age, is described. Although most of its appendages have been lost, preservation of several structures is vastly better than in most ‘orthodox’ fossils of these delicate animals. The well-preserved carapace suggests that, like extant representatives of these genera, but in no other daphniid, the animal exploited the surface film, beneath which it suspended itself by means of the straight ventral margins of its functionally bivalved carapace, each of which was armed with a row of close-set setae. Unlike the postabdomen of all the many extant anomopods, which is unsegmented, that of the fossil species, which was well preserved, was segmented; its distal half, or rather more, being clearly divided into four segments. Its topographically ventral margin was armed with pairs of very long spines (relatively longer than in any extant species), and the paired, hook-like terminal spines, clearly homologous with the claws of living species, were modified in a way that has no exact parallel in extant forms. The distal segment shows signs of annulations and may have had slight independent mobility. It is suggested that the long ventral spines of the postabdomen facilitated levering the animal forward over soft substrata. Its minute size reflects the fact that it is a first instar. Moreover, it was possibly squeezed out of the brood pouch prematurely when its parent became trapped in resin. The difficulty of assigning a place to the fossil in the phylogeny of the anomopods, all of which, including fossil forms of greater antiquity, have an unsegmented postabdomen, is briefly noted. The possibility that this attribute represents a very ancient atavism is tentatively suggested.     

Systematics and fossil evidence of host-parasitoid relationships of Calamiscus Borgmeier (Diptera: Phoridae)

The life history and hitherto unknown males of Calamiscus cryptopalpis are described. A second species of this genus, C. inflativenis n. sp. is described from Costa Rica. The species C. cryptopalpis is a gregarious parasitoid of injured stingless bees in Costa Rica. The co-occurrence of a specimen of the genus Calamiscus in a piece of amber with the oldest known fossil bee is hypothesized to be the most ancient host-parasitoid relationship between a pair of insect species documented thus far. The otherwise rarely-seen fly was probably lured to the amber site by the alarm pheromones of the bee, and then trapped in the sticky tree resins.     

A new genus and species of Cixiidae (Homoptera: Fulgoroidea) from Lower Cretaceous amber

Mundopoides aptianus gen. et sp. nov. (Homoptera: Fulgoroidea: Cixiidae) is described on the basis of a fossil of an adult female preserved in Lebanese amber of Lower Cretaceous (Aptian) age, and is compared with modern genera.     

A description of Siphonophora portoricensis Brandt (Diplopoda: Siphonophorida: Siphonophoridae), with a Catalogue of Ordinal Representatives in the New World

Data on geographical distribution are provided for S. portoricensis Brandt, the type species of Siphonophora Brandt and the oldest species-group name in the genus, family, and Order; it appears to occur throughout Puerto Rico. It is characterized by fusion of the distal three podomeres of the anterior gonopods, which are sublinear in shorter, less mature males and bend caudad for varying lengths in longer individuals. In the New World, the Siphonophorida is represented by both families, the Siphonorhinidae by one genus and one species, Illacme plenipes Cook and Loomis, from San Benito County, California, and the Siphonophoridae by 10 genera, six monotypic, 66 extant nominal species, and one subspecies; two fossil species of Siphonophora have been proposed along with an unnamed fossil form of Siphonocybe Pocock. The following new combinations are officially recorded: Columbianum gracilicornis (Carl), Columbiozonium pearsei (Chamberlin), Cordillerium fuhrmanni (Carl), Yucatanium sabachana (Chamberlin), and Guatemalium barberi (Chamberlin). The Siphonophoridae ranges from southern California, northcentral Arizona, and central Texas to the Andes of central Peru and São Paulo state, Brazil. It also occurs on 10 islands in the Greater and Lesser Antilles, being newly recorded from Martinique, but apparently does not cross the Straits of Florida and is unknown from Florida, the Bahamas, and the Turks and Caicos Islands.     

Retromalisus damzeni,gen. et sp. nov., a second Baltic amber taxon of the extinct family Berendtimiridae (Insecta: Coleoptera)

ABSTRACT

A second Baltic amber taxon, Retromalisus damzeni, gen. et sp. nov., is discovered in the previously monotypic extinct family Berendtimiridae. The morphological portrait of Berendtimiridae is complemented with the data on the structure of prosternum and abdomen.     

Some new African Ulopini (Homoptera,Cicadellidae, Ulopinae)

Summary

Two new genera and two new species of Ulopini are described from Africa; the distribution, and some aspects of the evolution, of genera in this tribe are discussed. It is suggested that Ulopa damaspina Dlabola belongs to the genus Moonia Distant and a new synonmy is suggested. In an Appendix the genera and species of the Ulopini are listed according to their geographical distribution.     

A new hyaliodine genus for Megacoelum nigroscutellatum Distant with new combinations and tribal reassignment for some other species of Miridae (Heteroptera)

Summary

A new genus is erected for Megacoelum nigroscutellatum Distant in the Hyaliodinae as opposed to the Mirinae where the species was originally described. Two species previously described in the genus, Araspus Distant, viz. A. scutellaris Poppius and A. bicolor Poppius, are considered not congeneric with the genotype of the genus and are therefore transferred tentatively to Imogen Kirkaldy in the Saturniomirini. The genus Attractotomelus recently erected by Linnavuori (1975) for Attractotomus serosus Odhiambo is synonymized with Millermiris Carvalho while the transfer of the latter to the Phylinae by Schuh (1974) is confirmed. Proboscidocoris nitidus Odhiambo is transferred to Polymerus Hahn while Polymerus capitatus (Distant) is transferred to Proboscidocoris Reuter and P. antennatus (Distant) to Charagochilus Fieber.     

First record of a non-pollinating fig wasp (Hymenoptera: Sycophaginae) from Dominican amber,with estimation of the size of its host figs

Fig trees and their pollinating fig wasps arose about 75 million years ago in the Cretaceous period. Several other groups of chalcid wasps also utilize figs for larval development, including sycophagines, the putative sister group to pollinating fig wasps. Whereas stone and amber fossil pollinators are known, no fossils representing non-pollinating fig wasp groups have been confirmed previously. Here, we describe the first Sycophaginae from the c.15–20 Ma Dominican amber, Idarnes thanatos sp. nov. Farache, Rasplus, Pereira and Compton, and discuss its relationships within the Idarnes carme species group. Additionally, we use linear regression to compare body size, ovipositor sheaths length, and host fig size data from extant Idarnes species to estimate the size of its host figs. Idarnes thanatos was most likely associated with small to medium sized figs (diameter ≤1.0 cm), that were likely to have been dispersed by birds and primates. The discovery of this close relative of extant non-pollinating fig wasps suggests that early Miocene and modern fig wasp communities may share similar ecological and functional features.     

Plantes Cryptogames de France. Par I. B. H. I. Desmazières. Fasc. XVII. Lille, 1836

Summary

A new genus and species of pilargid polychaete, Litocorsa stremma, is described from silty grounds on the west coast of Scotland. Its relationship to other pilargid genera is discussed. Notes on two other pilargid species and a revised key to the genera of the Pilargidae are given.